Articles | Volume 19, issue 11
https://doi.org/10.5194/gmd-19-4907-2026
https://doi.org/10.5194/gmd-19-4907-2026
Model description paper
 | 
12 Jun 2026
Model description paper |  | 12 Jun 2026

BORIS-2 – a benthic ecosystem model based on allometry

Adrian P. Martin, Anieke Brombacher, Noëlie Benoist, Brian J. Bett, Jennifer M. Durden, Sophy Oliver, and Andrew Yool

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Preprint under review for ESSD
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Cited articles

Atwood, T. B., Witt, A., Mayorga, J., Hammill, E., and Sala, E.: Global patterns in marine sediment carbon stocks, Front. Mar. Sci. ,7, 165, https://doi.org/10.3389/fmars.2020.00165, 2020. 
Benoist, N.: Advances in the state-of-the-art in the quantitative ecology of the marine megabenthos, PhD thesis, University of Southampton, 289 pp., https://eprints.soton.ac.uk/444730/1/Benoist_Noelie_PhD_thesis_Oct_2020.pdf (last access: 8 June 2026), 2020. 
Benoist, N. M. A., Bett, B. J., Morris, K. J., and Ruhl, H. A.: A generalised volumetric method to estimate the biomass of photographically surveyed benthic megafauna, Progress in Oceanography, 178, 102188, https://doi.org/10.1016/j.pocean.2019.102188, 2019. 
Best, M. A., Wither, A. W., and Coates, S.: Dissolved oxygen as a physico-chemical supporting element in the Water Framework Directive, Mar. Pollut. Bull., 55, 1–6, https://doi.org/10.1016/j.marpolbul.2006.08.037, 2007. 
Bett, B. J.: UK Atlantic Margin Environmental Survey: Introduction and overview of bathyal benthic ecology, Cont. Shelf Res., 21, 917–956, https://doi.org/10.1016/S0278-4343(00)00119-9, 2001. 
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Although under huge pressure and with permanent darkness, the seafloor has a diversity of organisms. They are almost entirely dependent on food sinking from the ocean surface onto the seafloor. This model allows us to study how these organisms survive in this hostile environment. Making use of evidence that biological characteristics, like lifetime, vary with size of organism and temperature, this model can simulate the fate of seafloor creatures from bacteria to large sea cucumbers.
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