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  <front>
    <journal-meta><journal-id journal-id-type="publisher">GMD</journal-id><journal-title-group>
    <journal-title>Geoscientific Model Development</journal-title>
    <abbrev-journal-title abbrev-type="publisher">GMD</abbrev-journal-title><abbrev-journal-title abbrev-type="nlm-ta">Geosci. Model Dev.</abbrev-journal-title>
  </journal-title-group><issn pub-type="epub">1991-9603</issn><publisher>
    <publisher-name>Copernicus Publications</publisher-name>
    <publisher-loc>Göttingen, Germany</publisher-loc>
  </publisher></journal-meta>
    <article-meta>
      <article-id pub-id-type="doi">10.5194/gmd-13-805-2020</article-id><title-group><article-title>Development and testing scenarios for implementing <?xmltex \hack{\break}?> land use and land cover changes during the Holocene <?xmltex \hack{\break}?> in Earth system model experiments</article-title><alt-title>Development and testing scenarios for implementing land use and land cover changes</alt-title>
      </title-group><?xmltex \runningtitle{Development and testing scenarios for implementing land use and land cover changes}?><?xmltex \runningauthor{S.~P.~Harrison et al.}?>
      <contrib-group>
        <contrib contrib-type="author" corresp="yes" rid="aff1">
          <name><surname>Harrison</surname><given-names>Sandy P.</given-names></name>
          <email>s.p.harrison@reading.ac.uk</email>
        <ext-link>https://orcid.org/0000-0001-5687-1903</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff2">
          <name><surname>Gaillard</surname><given-names>Marie-José</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff3 aff4">
          <name><surname>Stocker</surname><given-names>Benjamin D.</given-names></name>
          
        <ext-link>https://orcid.org/0000-0003-2697-9096</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff5">
          <name><surname>Vander Linden</surname><given-names>Marc</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff6 aff7">
          <name><surname>Klein Goldewijk</surname><given-names>Kees</given-names></name>
          
        <ext-link>https://orcid.org/0000-0003-2714-7507</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff8">
          <name><surname>Boles</surname><given-names>Oliver</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff9">
          <name><surname>Braconnot</surname><given-names>Pascale</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff10">
          <name><surname>Dawson</surname><given-names>Andria</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff11">
          <name><surname>Fluet-Chouinard</surname><given-names>Etienne</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff12 aff13">
          <name><surname>Kaplan</surname><given-names>Jed O.</given-names></name>
          
        <ext-link>https://orcid.org/0000-0001-9919-7613</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff14">
          <name><surname>Kastner</surname><given-names>Thomas</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff15">
          <name><surname>Pausata</surname><given-names>Francesco S. R.</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff16">
          <name><surname>Robinson</surname><given-names>Erick</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff17">
          <name><surname>Whitehouse</surname><given-names>Nicki J.</given-names></name>
          
        <ext-link>https://orcid.org/0000-0002-7044-6492</ext-link></contrib>
        <contrib contrib-type="author" corresp="no" rid="aff18 aff19 aff20">
          <name><surname>Madella</surname><given-names>Marco</given-names></name>
          
        </contrib>
        <contrib contrib-type="author" corresp="no" rid="aff8">
          <name><surname>Morrison</surname><given-names>Kathleen D.</given-names></name>
          
        </contrib>
        <aff id="aff1"><label>1</label><institution>Department of Geography and Environmental Science, University of Reading, Reading, UK</institution>
        </aff>
        <aff id="aff2"><label>2</label><institution>Department of Biology and Environmental Science, Linnaeus University, Kalmar, Sweden</institution>
        </aff>
        <aff id="aff3"><label>3</label><institution>Ecological and Forestry Applications Research Centre, Cerdanyola del Vallès, Spain</institution>
        </aff>
        <aff id="aff4"><label>4</label><institution>Department of Earth System Science, Stanford University, Stanford, CA 94305, USA</institution>
        </aff>
        <aff id="aff5"><label>5</label><institution>Department of Archaeology, University of Cambridge, Cambridge, UK</institution>
        </aff>
        <aff id="aff6"><label>6</label><institution>PBL Netherlands Environmental Assessment Agency, the Hague, the
Netherlands</institution>
        </aff>
        <aff id="aff7"><label>7</label><institution>Copernicus Institute of Sustainable Development, Utrecht University, Utrecht, the Netherlands</institution>
        </aff>
        <aff id="aff8"><label>8</label><institution>University Museum of Archaeology &amp; Anthropology, University of
Pennsylvania, Philadelphia, USA</institution>
        </aff>
        <aff id="aff9"><label>9</label><institution>Laboratoire des Sciences du Climat et de l'Environnement, Gif-sur-Yvette, France</institution>
        </aff>
        <aff id="aff10"><label>10</label><institution>Department of General Education, Mount Royal University, Calgary, Canada</institution>
        </aff>
        <aff id="aff11"><label>11</label><institution>Department of Earth System Science, Stanford University, Stanford, California, USA</institution>
        </aff>
        <aff id="aff12"><label>12</label><institution>Department of Earth Sciences, University of Hong Kong, Hong Kong</institution>
        </aff>
        <aff id="aff13"><label>13</label><institution>Institute of Geography, University of Augsburg, Augsburg, Germany</institution>
        </aff>
        <aff id="aff14"><label>14</label><institution>Senckenberg Biodiversity and Climate Research Centre, Frankfurt am Main, Germany</institution>
        </aff>
        <aff id="aff15"><label>15</label><institution>Centre ESCER, Department of Earth and Atmospheric Sciences, University of Quebec in Montreal, Montreal, Canada</institution>
        </aff>
        <aff id="aff16"><label>16</label><institution>Department of Anthropology, University of Wyoming, Laramie, Wyoming, USA</institution>
        </aff>
        <aff id="aff17"><label>17</label><institution>School of Geography, Earth and Environmental Science, University of Plymouth, Plymouth, UK</institution>
        </aff>
        <aff id="aff18"><label>18</label><institution>Department of Humanities (CaSEs), University Pompeu Fabra, Barcelona, Spain</institution>
        </aff>
        <aff id="aff19"><label>19</label><institution>ICREA Passeig Lluís Companys 23, 08010 Barcelona, Spain</institution>
        </aff>
        <aff id="aff20"><label>20</label><institution>School of Geography, Archaeology and Environmental Studies, University of the Witwatersrand, <?xmltex \hack{\break}?> Johannesburg, South Africa</institution>
        </aff>
      </contrib-group>
      <author-notes><corresp id="corr1">Sandy P. Harrison (s.p.harrison@reading.ac.uk)</corresp></author-notes><pub-date><day>2</day><month>March</month><year>2020</year></pub-date>
      
      <volume>13</volume>
      <issue>2</issue>
      <fpage>805</fpage><lpage>824</lpage>
      <history>
        <date date-type="received"><day>5</day><month>May</month><year>2019</year></date>
           <date date-type="rev-request"><day>23</day><month>July</month><year>2019</year></date>
           <date date-type="rev-recd"><day>12</day><month>January</month><year>2020</year></date>
           <date date-type="accepted"><day>20</day><month>January</month><year>2020</year></date>
      </history>
      <permissions>
        <copyright-statement>Copyright: © 2020 Sandy P. Harrison et al.</copyright-statement>
        <copyright-year>2020</copyright-year>
      <license license-type="open-access"><license-p>This work is licensed under the Creative Commons Attribution 4.0 International License. To view a copy of this licence, visit <ext-link ext-link-type="uri" xlink:href="https://creativecommons.org/licenses/by/4.0/">https://creativecommons.org/licenses/by/4.0/</ext-link></license-p></license></permissions><self-uri xlink:href="https://gmd.copernicus.org/articles/13/805/2020/gmd-13-805-2020.html">This article is available from https://gmd.copernicus.org/articles/13/805/2020/gmd-13-805-2020.html</self-uri><self-uri xlink:href="https://gmd.copernicus.org/articles/13/805/2020/gmd-13-805-2020.pdf">The full text article is available as a PDF file from https://gmd.copernicus.org/articles/13/805/2020/gmd-13-805-2020.pdf</self-uri>
      <abstract><title>Abstract</title>
    <p id="d1e357">Anthropogenic changes in land use and land cover (LULC) during the
pre-industrial Holocene could have affected regional and global climate.
Existing scenarios of LULC changes during the Holocene are based on
relatively simple assumptions and highly uncertain estimates of population
changes through time. Archaeological and palaeoenvironmental reconstructions have the potential to refine these assumptions and estimates. The Past Global Changes (PAGES) LandCover6k initiative is working towards improved reconstructions of LULC globally. In this paper, we document the types of archaeological data that are being collated and how they will be used to improve LULC reconstructions. Given the large methodological uncertainties involved, both in reconstructing LULC from the archaeological data and in implementing these reconstructions into global scenarios of LULC, we propose a protocol to evaluate<?pagebreak page806?> the revised scenarios using independent pollen-based reconstructions of land cover and climate. Further evaluation of the revised scenarios involves carbon cycle model simulations to determine whether the LULC reconstructions are consistent with constraints provided by ice core records of <inline-formula><mml:math id="M1" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> evolution and modern-day LULC. Finally, the protocol outlines how the improved LULC reconstructions will be used in palaeoclimate simulations in the Palaeoclimate Modelling Intercomparison Project to quantify the magnitude of anthropogenic impacts on climate through time and ultimately to improve the realism of Holocene climate simulations.</p>
  </abstract>
    </article-meta>
  </front>
<body>
      

<sec id="Ch1.S1" sec-type="intro">
  <label>1</label><title>Introduction and motivation</title>
      <p id="d1e380">Today, ca. 10 % of the ice-free land surface is estimated to be intensively
managed, and much of the reminder is under less intense anthropogenic use or
influenced by human activities (Arneth et al., 2019). Substantial
transformations of natural ecosystems by humans began with the geographically diachronous shift from hunting and gathering characteristic of the Mesolithic to cultivation and more permanent settlement during the Neolithic period (Mazoyer and Roudart, 2006; Zohary et al., 2012; Tauger, 2013; Maezumi et al., 2018), although there is controversy about the relative importance of climate changes and human impact on landscape development both during and since that time. Resolving the uncertainty about the extent and timing of land use is important because changes in land cover as a result of land use (land use land cover: LULC) have the potential to impact climate and the carbon cycle. Direct climate impacts occur through changes in the surface energy budget resulting from modifications of surface albedo,
evapotranspiration, and canopy structure (biophysical impacts; e.g. Pongratz
et al., 2010; Myhre et al., 2013; Perugini et al., 2017). LULC affects the
carbon cycle through modifications in vegetation, soil carbon storage
(biogeochemical impacts; e.g. Pongratz et al., 2010; Mahowald et al., 2017),
and turnover times, which change the C sink and source capacity of the
terrestrial biosphere. LULC changes have substantially contributed to the
increase in atmospheric greenhouse gases during the industrial period (Le Quéré et al., 2018). It has been suggested that greenhouse gas
emissions associated with Neolithic LULC changes were sufficiently large to
offset climate cooling after the mid-Holocene (the overdue glaciation hypothesis: Ruddiman, 2003). Although this has been challenged for several
reasons, including inconsistency with the land carbon balance derived from
ice core and peat records (e.g. Joos et al., 2004; Kaplan et al., 2011;
Singarayer et al., 2011; Mitchell et al., 2013; Stocker et al., 2017), an LULC
impact on climate in more recent millennia appears more plausible.</p>
      <p id="d1e383">Climate model simulations have shown that LULC changes have discernible impacts on climate, both in regions with large prescribed changes in LULC
and in teleconnected regions with no major local human activity (Vavrus et
al., 2008; Pongratz et al., 2010; He et al., 2014; Smith et al., 2016). At
the global scale, the biogeophysical effects of the accumulated LULC change
during the Holocene, which resulted in reconstructed land cover patterns in
1850 CE, have been estimated to cause a slight cooling (0.17 <inline-formula><mml:math id="M2" display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula>C)
that is offset by biogeochemical warming (0.9 <inline-formula><mml:math id="M3" display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula>C), giving a
net global warming (0.73 <inline-formula><mml:math id="M4" display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula>C) (He et al., 2014). However, in these
simulations, biophysical and biogeochemical effects were of comparable
magnitude in the most intensively altered landscapes of Europe, Asia, and
North America (He et al., 2014). Using parallel simulations with and
without LULC changes, Smith et al. (2016) showed that detectable temperature
changes due to LULC could have occurred as early as 7000 years ago (7 ka BP) in summer and throughout the year by 3 ka BP. All of these conclusions, however, are obviously contingent on the imposed LULC forcing, which is highly uncertain.</p>
      <p id="d1e413">There have been several attempts to map LULC changes through time (e.g. Ramankutty and Foley, 1999; Pongratz et al., 2008; Kaplan et al., 2011;
Klein Goldewijk et al., 2011, 2017a, b). All of these reconstructions assume that anthropogenic land use is a function of population density and the suitability of land for crops and/or pasture. Estimates of regional population trends are then used through time in combination with assumptions about per-capita land use and spatial land use allocation schemes to estimate anthropogenic changes in LULC across time and space. However, differences in the underlying assumptions about land use per capita, which are generalized from limited and often site-specific data, have resulted in large differences in the final reconstructions (Gaillard et al., 2010; Kaplan et al., 2017). Hence, there are still very large uncertainties about the timing and magnitude of LULC changes, both at a global and at a regional scale (Fig. 1).</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F1"><?xmltex \currentcnt{1}?><label>Figure 1</label><caption><p id="d1e419">Land use at ca. 6000 years ago (6 ka BP, 4000 years BCE) from the
two widely used global historical land use scenarios HYDE 3.2 (<bold>a</bold>;
Klein Goldewijk et al., 2017a) and KK10 (<bold>b</bold>; Kaplan et al., 2011),
illustrating the large disagreement between LULC scenarios at a regional scale. In both scenarios, the land–sea mask and lake areas are for the present day.</p></caption>
        <?xmltex \igopts{width=236.157874pt}?><graphic xlink:href="https://gmd.copernicus.org/articles/13/805/2020/gmd-13-805-2020-f01.png"/>

      </fig>

      <p id="d1e434">There is a wealth of archaeological, historical, and palaeo-vegetation data
that could be used to improve the relatively simple rules used to generate
global LULC reconstructions. For example, settlement density and numbers of
radiocarbon-dated artefacts can be used to infer population sizes and their
temporal dynamics (Rick, 1987; Williams, 2012; Silva and Vander Linden, 2017). Carbonized and waterlogged plant remains and animal bones can be used
to infer the occurrence and nature of agriculture at a site, although their
presence provides no quantitative information about the area under
cultivation (Wright, 2003; Lyman, 2008; Orton et al., 2016). Although the
record of LULC is likely to be patchy and incomplete because of preservation and sampling issues, the systematic use of archaeological data is one important way to improve current LULC scenarios.</p>
      <p id="d1e437">The Past Global Change (PAGES; <uri>http://www.pastglobalchanges.org/</uri>, last access: 14 February 2020) LandCover6k Working Group
(<uri>http://pastglobalchanges.org/ini/wg/landcover6k</uri>, last access: 14 February 2020) is currently working to develop a rigorous and robust approach to provide data and data products that can be used to inform<?pagebreak page807?> the development of LULC scenarios (Gaillard et al., 2018). LULC changes are taken into account in climate model simulations currently being made in the current phase of the Coupled Model Intercomparison Project (CMIP6) for the historic period and the future scenario runs (Eyring et al., 2016). They are also included in climate model simulations of the past millennium (Jungclaus et al., 2017) in order to ensure that these runs mesh seamlessly with the historic simulations. However, the Land Use Harmonization dataset (LUH2: Hurtt et al., 2017) only extends back to 850 CE, and thus scenarios of LULC changes are currently not included in the CMIP6 palaeoclimate simulations, including mid-Holocene simulations, that are used as a test of how well state-of-the-art climate models reproduce large climate changes. In this paper, we discuss how archaeological data will be used to improve global LULC scenarios for the Holocene. Given that there are large uncertainties associated with the primary data and further uncertainties may be introduced when this information is used to modify existing LULC scenarios, we outline a series of tests that will be used to evaluate whether the revised LULC scenarios are consistent with the changes implied by independent pollen-based
reconstructions of land cover and whether they produce more realistic
estimates of both carbon cycle and climate changes. Finally, we present a
protocol for implementing LULC in Earth system model simulations to be
carried out in the current phase of the Palaeoclimate Modelling
Intercomparison Project (PMIP: Otto-Bleisner et al., 2017; Kageyama et al.,
2018). However, the datasets and protocol will also be useful in later
phases of other CMIP projects, including the Land Use Model Intercomparison
Project (LUMIP) and the Land Surface, Snow and Soil Moisture Model
Intercomparison Project (LS3MIP) (Lawrence et al., 2016; van den Hurk et al., 2016).</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F2" specific-use="star"><?xmltex \currentcnt{2}?><label>Figure 2</label><caption><p id="d1e448">Proposed scheme for developing robust LULC scenarios through iterative testing and refinement as input to Earth system model (ESM)
simulations. The archaeological inputs developed in phase 1 can be used
independently or together to improve the LULC reconstructions; iterative
testing of the LULC scenario reconstruction (phase 2) will ensure that these
inputs are reliable before they are used for ESM simulations (phase 3). The
uppermost three LULC simulations capitalize on already planned baseline
simulations without LULC; the lowermost two simulations are envisaged as new
sensitivity experiments.</p></caption>
        <?xmltex \igopts{width=341.433071pt}?><graphic xlink:href="https://gmd.copernicus.org/articles/13/805/2020/gmd-13-805-2020-f02.png"/>

      </fig>

</sec>
<sec id="Ch1.S2">
  <label>2</label><title>LandCover6k methodology</title>
      <p id="d1e465">The primary source of information about human exploitation of the landscape
comes from archaeological data. In general, these data are site specific,
spatiotemporal coverage is often patchy, and the types and quality of
evidence available vary between sites and regions. Generalizing from site-specific data to landscape or regional scales involves making
assumptions about human behaviour and cultural practices. Because of the
inherent uncertainties, we advocate an iterative approach to incorporate
archaeological data into LULC scenarios in LandCover6k (Fig. 2). We propose
revising the existing LULC scenarios through the incorporation of diverse
archaeological inputs (Fig. 2, phase 1; see Sects. 3 and 4) and testing
the revised LULC scenarios for their plausibility and consistency with other
lines of evidence (Fig. 2, phase 2 with iterative testing; see Sects. 5–7). As a first test, the revised LULC scenarios of the extent of cropland and grazing land through time will be compared with independent data on land cover changes, specifically pollen-based reconstructions of the extent of open land (see e.g. Trondman et al., 2015; Kaplan et al., 2017) (Sect. 5). Further testing the LULC scenarios involves sensitivity tests using global climate models (Sect. 6) and global carbon cycle models (Sect. 7). While the computational cost of the climate model simulations can be minimized using equilibrium time-slice simulations, the carbon cycle constraint relies on transient simulations but may be derived from uncoupled, land-only simulations. Simulated climates at key times can be evaluated against reconstructions of climate variables (e.g. Bartlein et al., 2011) (Sect. 6). The parallel evolution of <inline-formula><mml:math id="M5" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> and its isotopic composition (<inline-formula><mml:math id="M6" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">13</mml:mn></mml:msup><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula>) can be used to derive the carbon balance of
the terrestrial biosphere and the ocean separately (Elsig et al., 2009);
in combination with estimates for other contributors to land carbon changes
such as C sequestration by peat build-up, it provides a strong constraint on the
evolution of LULC through time. An underprediction or overprediction of anthropogenic LULC-related <inline-formula><mml:math id="M7" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> emissions during a specific interval results in consequences for the dynamics of the atmospheric greenhouse gas burden in subsequent times (Stocker et al., 2017) (Sect. 7). Thus, these tests can be used to identify issues in the original archaeological datasets and/or the way these data were incorporated into the LULC scenarios that require further refinement. Phase 3 of the project (Fig. 2) provides a protocol for the<?pagebreak page808?> implementation of the revised LULC scenarios in Earth system model simulations (Sect. 8).</p>
</sec>
<sec id="Ch1.S3">
  <label>3</label><title>Archaeological data inputs</title>
      <p id="d1e511">LandCover6k is creating a number of products that will be used to improve
the LULC scenarios (Fig. 2). Here, we summarize the important features of
these data products before showing how they will be incorporated within a
scenario development framework.</p>
<sec id="Ch1.S3.SS1">
  <label>3.1</label><?xmltex \opttitle{Population dynamics from {$\protect\chem{{}^{{14}}C}$}~data}?><title>Population dynamics from <inline-formula><mml:math id="M8" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi/><mml:mn mathvariant="normal">14</mml:mn></mml:msup><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula> data</title>
      <p id="d1e534">Radiocarbon is the most routinely used absolute dating technique in archaeology, especially for the Holocene. Many thousands of radiocarbon
dates are available from the archaeological literature. A number of regional
and pan-regional initiatives are compiling these records through exhaustive
surveys of the archaeological literature (e.g. the Canadian Archaeological
Radiocarbon Database: <uri>https://www.canadianarchaeology.ca/</uri>, last access: 14 February 2020). Statistical approaches, such as summed probability distributions (SPDs), can then be used to infer past demographic fluctuations from these compilations (Fig. 3). This method assumes that the more people there were, the more remains of their various activities they left behind and that this is directly reflected in the number of samples excavated and dated (Rick, 1987: Robinson et al., 2019). There are biases that could affect the expected one-to-one relationship between the number of people and the number of radiocarbon dates on archaeological material, including lack of uniform sampling through time and space caused by different archaeological research interests and traditions in different regions as well as increased preservation issues with increasing age, but these can be minimized by auditing the datasets. Assessment of the robustness of population reconstructions through time can be made statistically by comparing a null hypothesis of demographic growth constructed from an exponential fit to the data with the actual record of the number of dates through time (Shennan et al., 2013; Timpson et al., 2014). Mathematical simulations show that the method is relatively robust for large sample sizes (Williams, 2012). Radiocarbon dates have been successfully used in several regions to identify population fluctuations associated with hunter–gatherers (Japan: Crema et al., 2016) and the introduction of farming and subsequent changes in farming regimes (western Europe: Shennan et al., 2013; Wyoming: Zahid et al., 2016; South Korea: Oh et al., 2017; see also Freeman et al., 2018) as well as climatic oscillations (Ireland: Whitehouse et al., 2014).</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F3" specific-use="star"><?xmltex \currentcnt{3}?><label>Figure 3</label><caption><p id="d1e542">Reconstruction of changes in population size on the Iberian Peninsula during the Holocene (9000 to 2000 BP, 9 ka to 2 ka BP) using summed probability distributions (SPDs) of radiocarbon dates <bold>(a)</bold> (data after Balsera et al., 2015). The red line indicates the onset of agriculture in the region. Panels <bold>(b)</bold> and <bold>(c)</bold> show areas under human use at 6 ka <bold>(b)</bold> and 4 ka <bold>(c)</bold> using kernel density estimates; the white dots are actual archaeological sites, and the shading shows the implied density of occupation.</p></caption>
          <?xmltex \igopts{width=341.433071pt}?><graphic xlink:href="https://gmd.copernicus.org/articles/13/805/2020/gmd-13-805-2020-f03.png"/>

        </fig>

</sec>
<?pagebreak page809?><sec id="Ch1.S3.SS2">
  <label>3.2</label><title>Date of first agriculture</title>
      <p id="d1e574">Radiocarbon dates can also be used to track the timing and process of dispersal events, such as the diffusion of plant and animal domesticates
from their initial centres of domestication. Since the distribution of samples is often patchy, geostatistical techniques such as kriging and
splines are used to spatially interpolate the information in order to
provide quantitative estimates of the timing of spread. Work carried out in
Europe (Bocquet-Appel et al., 2009), Asia (Silva et al., 2015), and Africa
(Russell et al., 2014) demonstrates that there are different rates of diffusion even within a region, reflecting the possible impact of natural
features (e.g. waterways, elevation, ecology) on diffusion rates (Davison et
al., 2006; Silva and Steele, 2014). Numerous studies provide robust local
estimates for the earliest regional occurrence of agriculture, and these are
being synthesized to provide a global product within LandCover6k (Fig. 2).</p>
</sec>
<sec id="Ch1.S3.SS3">
  <label>3.3</label><title>Global land use and livestock maps</title>
      <p id="d1e585">Maps of the distribution of archaeological sites or of areas linked to a
given food production system have been produced for individual site catchments or small regions (e.g. Zimmermann et al., 2009; Barton et al.,
2010; Kay et al., 2019). LandCover6k is developing global land use maps for
specific time windows using a global hierarchical classification of land use categories (Morrison et al., 2018) based on land use types that are widely recognized from the archaeological record. At the highest level, the maps distinguish between areas where there is no (or only limited) evidence of land use and areas characterized by hunting, foraging, and fishing activities,
pastoralism, agriculture, and urban and/or extractive land use (Fig. 4). Except in
the cases in which land use is minimal (no human land use, extensive or minimal
land use), further distinctions are subsequently made to encompass the
diversity of land use activities in each land use type (Fig. 4). A third
level of distinction is made in the case of two categories (agroforestry,
wet cultivation) in which there are very different levels of intervention in
different regions. Explanations of this terminology are given in Morrison et
al. (2018). The LandCover6k land use maps (see e.g. Fig. 5) will be based on
different methods ranging from kernel density estimates to expert
assessments depending on the quality and quantity of the archaeological
information available from different regions.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F4" specific-use="star"><?xmltex \currentcnt{4}?><label>Figure 4</label><caption><p id="d1e590">The hierarchical scheme of land use classes used for global mapping in LandCover6k (updated from Morrison et al., 2018).</p></caption>
          <?xmltex \igopts{width=398.338583pt}?><graphic xlink:href="https://gmd.copernicus.org/articles/13/805/2020/gmd-13-805-2020-f04.png"/>

        </fig>

      <?xmltex \floatpos{t}?><fig id="Ch1.F5"><?xmltex \currentcnt{5}?><label>Figure 5</label><caption><p id="d1e601">An example of regional land use mapping. <bold>(a, b)</bold> The
distribution of known archaeological sites superimposed on kernel density
estimates of the extent of land use based on the density of observations
and <bold>(c, d)</bold> these data superimposed on the LandCover6k
land use classes for the middle Neolithic (3600–3400 years BCE, 5600–5400 years BP, 5.6–5.4 ka BP) <bold>(a, c)</bold> and the early Neolithic (3750–3600 years BCE, 5750–5600 years BP, 5.7–5.6 ka BP) <bold>(b, d)</bold> in Ireland. Data points derive from <inline-formula><mml:math id="M9" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi/><mml:mn mathvariant="normal">14</mml:mn></mml:msup><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula>-dated archaeological sites and distributions of settlements and monuments that have been assigned to each archaeological period following the dataset published in McLaughlin et al. (2016). The assigned land use classes are inferred from archaeological material from one site (or multiple sites) within the grid box. It should not be assumed that the whole grid cell was being used for agriculture during the middle and early Neolithic. Informed assessment suggests that agricultural land (crop growing and grazing combined) probably occupied 10 %–15 % of the total grid area in the low-level food production regions of the eastern and western coastal areas, whilst agricultural land likely represents 5 % or less of the total grid cell area in inland areas.</p></caption>
          <?xmltex \igopts{width=236.157874pt}?><graphic xlink:href="https://gmd.copernicus.org/articles/13/805/2020/gmd-13-805-2020-f05.png"/>

        </fig>

      <p id="d1e635">There is considerable variation in how intensely land is used both for crops
and for grazing within broad land use categories both geographically and
through time (Ford and Clarke, 2015; Styring et al., 2017). Maps of land use
types do not provide direct information on the intensity of farming practices or how they translate into per-capita land use. Archaeological data about agricultural yields, combined with information from analogous contemporary cultures, historical information (e.g. Pongratz et al., 2008), and theoretical estimates of land use required to meet dietary and energy requirements (e.g. Hughes et al., 2018), can be used to provide regional estimates of per-capita land use for specific land use categories. LandCover6k will synthesize this information to<?pagebreak page810?> allow regionally specific estimates of per-capita land use to be derived from the global land use maps.</p>
      <p id="d1e638">Information about the extent of grazing land is an important input for the
development of revised LULC scenarios but from a carbon cycle modelling
perspective, the amount of biomass removed by grazing is also a key parameter. Biomass loss varies not only with population size but also with
the type of animal being reared (Herrero et al., 2013; Phelps and Kaplan, 2017), and thus information about what animals were present at a given
location and estimates of population sizes are needed for improving the
existing LULC scenarios. Although the conditions of bone preservation vary
across the globe due to factors such as soil acidity, animal bones are routinely excavated (Lyman, 2008; Reitz and Wing, 2008). Morphometric
analyses of bones, along with collateral information such as age-related
culling patterns, make it possible to determine whether these are the remains of domesticated species. We thus have a relatively precise idea of when livestock were introduced into a region, what types of animals were being reared at a given time, and can also make informed estimates of population size. Although the level of detail will vary geographically, this information can be used to produce global livestock maps.</p>
      <p id="d1e641">The harvesting of wood for domestic fires, building, and industrial
activities such as transportation, pottery making, and metallurgy is an
important aspect of human exploitation of the landscape in the pre-industrial period (McGrath et al., 2015). It has been argued that even
Mesolithic hunter–gatherer communities shaped their environment through wood
harvesting (Bishop et al., 2015). Approaches have been developed to quantifying the wood harvest associated with archaeological settlements at
specific times based on the evidence of types of wood use, household energy
requirements, population size, and calorific value of the wood used (see
e.g. Marston, 2009; Janssen et al., 2017). However, quantitative information
on ancient technology and lifestyle is sparse, and direct estimates of the
amount of wood harvest through time are likely to remain highly uncertain
(Marston et al., 2017; Veal, 2017). Nevertheless, combining evidence-based
inferential approaches with improved estimates of population size should allow changes in wood harvesting to be taken into account in constructing
revised LULC scenarios.</p><?xmltex \hack{\newpage}?>
</sec>
</sec>
<?pagebreak page811?><sec id="Ch1.S4">
  <label>4</label><title>Incorporation of archaeological data in LULC scenarios</title>
      <p id="d1e654">The existing LULC scenarios are substantially dependent on historical regional population estimates at key times, which are then linearly
interpolated to provide a year-by-year estimate of population. Estimates of
regional population growth based on suitably screened <inline-formula><mml:math id="M10" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi/><mml:mn mathvariant="normal">14</mml:mn></mml:msup><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula> data can be used to modify existing population growth curves (Fig. 6), both in terms
of establishing the initial date of human presence and by modifying a linear
growth curve to allow for intervals of population growth and decline.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F6" specific-use="star"><?xmltex \currentcnt{6}?><label>Figure 6</label><caption><p id="d1e671">Schematic illustration of the proposed implementation of
<inline-formula><mml:math id="M11" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi/><mml:mn mathvariant="normal">14</mml:mn></mml:msup></mml:mrow></mml:math></inline-formula>C-based population estimates, date of first agriculture, land use
maps, and land use per-capita information in the HYDE model (here indicated
as HYDE3.x). The archaeological data are represented as values for a grid
cell in geographic space at a given time for date of first agriculture and
land use, but as a time series for a specific grid cell for population and
land use per capita. In the case of population estimates, date of first
agriculture, and land use per-capita data, we show the initial estimate and
the revised estimate after taking the archaeological information into
account in the HYDE3.x plot. It should be assumed in the case of the
land use mapping that the original estimate indicated no land use
in this region.</p></caption>
        <?xmltex \igopts{width=341.433071pt}?><graphic xlink:href="https://gmd.copernicus.org/articles/13/805/2020/gmd-13-805-2020-f06.png"/>

      </fig>

      <p id="d1e690">Information on the timing of the first appearance of agriculture at specific
locations can be used to constrain the temporal record of LULC changes in the scenarios. This information can also be used to allocate LULC changes
geographically across regions (Fig. 6). Global land use maps can be used
to identify areas where there was no permanent agricultural activity at a
given time (e.g. either unsettled areas or areas occupied by hunter–gatherer
communities) and provide a further constraint on the geographic extent of
the LULC changes given by the scenarios (Fig. 6). The type of agriculture,
including whether the region was predominantly used for tree or annual crops
or for pasture, modifies the area of open land specified in the LULC scenarios. Information on the extent of rainfed versus irrigated agriculture, as indicated by the presence of irrigation structures associated with archaeological sites, can also be used to refine the distribution of these classes in the LULC scenarios. Per-capita land use estimates and their changes through time (see e.g. Hughes et al., 2018; Weiberg et al., 2019) provide a further refinement of the LULC scenarios, allowing for a better characterization of the distinction between e.g. areas given over to extensive versus intensive animal production (rangeland versus pasture in the HYDE 3.2 terminology). There will remain areas of the world for which this kind of fine-grained information is not available. Nevertheless, by incorporating information when it exists, the LandCover6k products will contribute to a systematic refinement of existing LULC scenarios. Iterative testing of the revised scenarios will ensure that they are robust.</p>
</sec>
<sec id="Ch1.S5">
  <label>5</label><title>Using pollen-based reconstructions of land cover changes to evaluate LULC scenarios</title>
      <p id="d1e702">Pollen-based vegetation reconstructions can be used to corroborate archaeological information on the date of first agriculture from the
appearance of cereals and agricultural weeds. These reconstructions can also
be used to test the LULC reconstructions, either using relative changes in
forest cover or reconstructions of the area occupied by different land cover
types. LandCover6k uses the REVEALS pollen source-area model (Sugita, 2007)
to estimate vegetation cover from fossil pollen assemblages. REVEALS predicts the relationship between pollen deposition in large lakes and the abundance of individual plant taxa in the surrounding vegetation at a large spatial scale (ca. 100 km <inline-formula><mml:math id="M12" display="inline"><mml:mo>×</mml:mo></mml:math></inline-formula> 100 km; Hellman et al., 2008a, b) using models
of pollen dispersal and deposition. REVEALS can also be used with pollen
records from multiple small lakes or peat bogs (Trondman et al., 2016), although this results in larger uncertainties in the estimated area occupied
by individual taxa. The estimates obtained for individual taxa are summed to
produce estimates of the area occupied by either plant functional (e.g. summer-green trees, evergreen trees) or land cover (e.g. open land, grazing land, cropland) types.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F7" specific-use="star"><?xmltex \currentcnt{7}?><label>Figure 7</label><caption><p id="d1e714">Northern extratropical (<inline-formula><mml:math id="M13" display="inline"><mml:mrow><mml:mo>&gt;</mml:mo><mml:mn mathvariant="normal">40</mml:mn></mml:mrow></mml:math></inline-formula> <inline-formula><mml:math id="M14" display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula>N) mean fractional cover of open land at 6000 years ago (6 ka BP: <bold>b</bold>) and 200 years ago (0.2 ka BP: <bold>a</bold>) estimated using REVEALS and the difference in fractional cover between the two periods <bold>(c)</bold>. Red indicates an increase in open land and blue a decrease (after Dawson et al., 2018).</p></caption>
        <?xmltex \igopts{width=341.433071pt}?><graphic xlink:href="https://gmd.copernicus.org/articles/13/805/2020/gmd-13-805-2020-f07.png"/>

      </fig>

      <p id="d1e751">The geographic distribution of pollen records is uneven. There are also many
areas of the world where environments that preserve pollen (i.e. lakes, bogs, forest hollows) are sparse. Site-based reconstructions of land cover are therefore interpolated statistically to produce spatially continuous
reconstructions (Nielsen et al., 2012; Pirzamanbein et al., 2014, 2018). LandCover6k uses a 1<inline-formula><mml:math id="M15" display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> resolution grid and all available pollen records in each grid cell to produce an estimate of land cover per grid cell through time. The more pollen records per grid cell and pollen counts per time window, the smaller the estimated error on the land cover reconstruction. The uncertainties on the pollen-based REVEALS reconstructions are partly expressed by their standard errors (SEs). These SEs take into account the SE on the relative pollen productivity (RPP) of each plant taxon included in the REVEALS reconstructions and the variability between the site-specific REVEALS reconstructions (e.g. Trondman<?pagebreak page813?> et al., 2015). The uncertainties on the pollen-based land cover reconstructions are taken into account when these reconstructions are compared with LULC scenarios (Kaplan et al., 2017).</p>
      <p id="d1e764">The REVEALS approach has already been used to produce gridded reconstructions of changes in the amount of open land through time across the northern extratropics (Fig. 7; Dawson et al., 2018) These reconstructions provide mean plant cover for time slices of 500 years through the Holocene until 0.7 ka BP and three historical time windows (modern – 0.1, 0.1–0.35, and 0.35–0.7 ka BP). The more pollen samples per time interval and pollen records per grid cell, the more years within the 500-year time slice will be represented in the reconstruction. This implies that the number of years represented in a time-slice reconstruction varies in space and time.</p>
      <p id="d1e767">A major limitation in applying REVEALS globally is the requirement for
information about the relative pollen productivity (RPP) of individual pollen taxa, which is currently largely lacking for the tropics. However, LandCover6k has been collecting RPPs for China, south-east India, Cameroon, Brazil, and Argentina, and pollen-based land cover reconstructions will be
available for sufficient parts of the tropics to allow the testing of the LULC
scenarios. Another limitation of the REVEALS reconstructions is that RPP
estimates are available for cultivated cereals but not for other cultivars
or cropland weeds, so the LandCover6k pollen-based reconstructions will
generally underestimate cropland cover (Trondman et al., 2015). It may also
be possible to use alternative pollen-based reconstructions of land cover
changes, such as the modern analogue approach (MAT: e.g. Tarasov et al.,
2007; Zanon et al., 2018), pseudo-biomization (e.g. Fyfe et al., 2014), or
STEPPS (Dawson et al., 2016). While none of these methods require RPPs, MAT
and STEPPS can only be applied in regions where the pollen datasets have
dense coverage (such as Europe and North America), and pseudo-biomization is
affected by the non-linearity of the pollen–vegetation relationship that the
REVEALS approach is designed to remove.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F8" specific-use="star"><?xmltex \currentcnt{8}?><label>Figure 8</label><caption><p id="d1e772">Quantitative comparison of the change in climate between the
mid-Holocene (6 ka) and the pre-industrial period as shown by pollen-based
reconstructions gridded to <inline-formula><mml:math id="M16" display="inline"><mml:mrow><mml:mn mathvariant="normal">2</mml:mn><mml:mo>×</mml:mo><mml:mn mathvariant="normal">2</mml:mn></mml:mrow></mml:math></inline-formula><inline-formula><mml:math id="M17" display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> resolution to be compatible with the model resolution (from Bartlein et al., 2011) and in simulations with and without the incorporation of land use change (from Smith et al., 2016). This figure illustrates the approach that will be taken to evaluate the impact of new LULC scenarios on climate. The imposed land use changes at
6000 years ago (6 ka BP) were derived from the KK10 scenario (Kaplan et al., 2011). The plots show comparisons of mean annual temperature (MAT), mean
temperature of the coldest month (MTCO), and mean annual precipitation (MAP)
for the northern extratropics (north of 30<inline-formula><mml:math id="M18" display="inline"><mml:msup><mml:mi/><mml:mo>∘</mml:mo></mml:msup></mml:math></inline-formula> N); each dot
represents a model grid cell in which comparisons with the pollen-based
reconstructions are possible. Although the incorporation of land use produces
somewhat warmer and wetter climates in these simulations, overall the
incorporation of land use produces no improvement of the simulated climates
at sites with pollen-based reconstructions.</p></caption>
        <?xmltex \igopts{width=497.923228pt}?><graphic xlink:href="https://gmd.copernicus.org/articles/13/805/2020/gmd-13-805-2020-f08.png"/>

      </fig>

      <p id="d1e810">Comparison of the reconstructions of the extent of open land with the LULC
deforestation scenarios will provide a first evaluation of the realism of the revised LULC scenarios (e.g. Kaplan et al., 2017). Underestimation or
overestimation of open land in the LULC scenarios is not necessarily an indication that these scenarios are inaccurate because (a) pollen-based
reconstructions cannot distinguish between anthropogenic and climatically
determined natural open land (e.g. natural grasslands, steppes, wetlands), and<?pagebreak page814?> (b) REVEALS underestimates cropland cover because there are no RPP estimates for cultivars other than cereals. However, overestimation of the area of open land in the LULC scenarios might suggest problems either in the archaeological inputs or their implementation, especially for times or regions for which other evidence indicates cereals were the major crop. In this sense, despite potential problems, the LandCover6k pollen-based reconstructions of land cover will provide an important independent test of
the revised LULC scenarios.</p>
</sec>
<sec id="Ch1.S6">
  <label>6</label><title>Testing the reliability of improved scenarios using climate model
simulations</title>
      <p id="d1e821">A second test of the realism of the improved LULC scenarios is to examine
whether incorporating LULC changes improves the realism of the simulated
climate when compared to palaeoclimate reconstructions (Fig. 8). The
mid-Holocene (6000 years ago, 6 ka BP) is an ideal candidate for such a test because benchmark datasets of quantitative climate reconstructions are
available (e.g. Bartlein et al., 2011), the interval has been a focus
through multiple phases of PMIP, control simulations with no LULC have already been run, and evaluation of these simulations has identified regions
where there are major discrepancies between simulated and reconstructed
climates e.g. the observed expansion of Northern Hemisphere monsoons, climate changes over Europe, the magnitude of high-latitude warming, and wetter conditions in central Eurasia (Mauri et al., 2014; Harrison et al., 2015; Bartlein et al., 2017). There are discernible anthropogenic impacts on the landscape in many of these regions by 6 ka, although they are not as strong as during the later Holocene and they are not present everywhere. Nevertheless, the 6 ka BP interval provides a good focus for testing whether
improvements to the LULC scenarios produces more realistic simulations of climate. Such an evaluation would need to go beyond the global comparison
made here (Fig. 8) to regional comparisons to identify whether improvements in simulated climate in regions where there is a large anthropogenic impact on land cover result in a degradation in the simulated climate elsewhere.</p>
</sec>
<sec id="Ch1.S7">
  <label>7</label><title>Testing the reliability of improved scenarios using carbon cycle models</title>
      <p id="d1e832">Carbon cycle modelling will be used as a further test of the realism of the
improved LULC scenarios. Two constraints are available for testing the
realism of past LULC scenarios. First, reconstructions of LULC history must
converge on the present-day state, which is relatively well constrained by
satellite land cover observations and national statistics on the amount of
land under use. Reconstructing the extent of past LULC change thus reduces
to allocating a fixed total amount of land conversion from natural to
agricultural use over time. More conversion in earlier periods implies less
conversion in later periods. At the continental to global scale,<?pagebreak page815?> cumulative
LULC emissions scale linearly with the agricultural area. LULC scenarios
that converge to the present-day state also converge to within a small range
of cumulative historical emissions (Stocker et al., 2011, 2017). Deviations from a linear relationship between extent and emissions are due to differences in biomass density in potential natural and agricultural vegetation of different regions affected by anthropogenic LULC. Differences in cumulative emissions for alternative LULC reconstructions with an identical present-day state are due to the long response time of soil carbon content following a change in carbon inputs and soil cultivation. Conserving the total extent of LULC (and allocating a fixed total expansion over time) is thus approximately equivalent to conserving cumulative historical LULC emissions. Thus, more LULC <inline-formula><mml:math id="M19" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> emissions in earlier periods imply less <inline-formula><mml:math id="M20" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> emissions in more recent periods.</p>
      <p id="d1e857">The total C budget of the terrestrial biosphere provides a second constraint
on LULC emissions through time. The net C balance of the land biosphere, which reflects the sum of all natural and anthropogenic effects on terrestrial C storage, can be reconstructed from ice core data of past
<inline-formula><mml:math id="M21" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> concentrations and <inline-formula><mml:math id="M22" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">13</mml:mn></mml:msup><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula> composition (Elsig et al., 2009). Provided that all of the natural contributions to the land C
inventory (e.g. the build-up of natural peatlands: Loisel et al., 2014) can be specified from independent evidence, the anthropogenic sources can be estimated as the difference between the total terrestrial C budget and natural contributions (Fig. 9) at any specific time.</p>

      <?xmltex \floatpos{t}?><fig id="Ch1.F9"><?xmltex \currentcnt{9}?><label>Figure 9</label><caption><p id="d1e886">Illustration of the terrestrial C budget approach to evaluate LULC. The total terrestrial C balance (green circle, “total”) is constrained by ice core records of <inline-formula><mml:math id="M23" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> and its isotopic signature (<inline-formula><mml:math id="M24" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi mathvariant="italic">δ</mml:mi><mml:mn mathvariant="normal">13</mml:mn></mml:msup><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula>). Estimates for C balance changes in different natural land carbon cycle components (e.g. peatlands, permafrost, forest
expansion or retreat, desert greening) are estimated independently (blue
slices, “natural components”) either from empirical upscaling of site-scale
observations or from model-based analyses (carbon cycle models forced with varying climate). The remainder (yellow slice, “remainder”) is then calculated as the total terrestrial C balance (green circle, “total”) minus the sum of the separate estimates of the natural components (blue slices, “natural components”). The remainder is effectively the emissions resulting from LULC changes and can therefore be compared to LULC <inline-formula><mml:math id="M25" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> emission estimates by carbon cycle models.</p></caption>
        <?xmltex \igopts{width=199.169291pt}?><graphic xlink:href="https://gmd.copernicus.org/articles/13/805/2020/gmd-13-805-2020-f09.png"/>

      </fig>

      <p id="d1e931">Transient simulations with a model that simulates <inline-formula><mml:math id="M26" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> emissions in
response to anthropogenic LULC can be used to test the reliability of the
LULC scenarios by comparing results obtained with prescribed LULC changes
through time against a baseline simulation without imposed LULC. This will
necessitate making informed decisions about the fraction of land under
cultivation that is abandoned or left fallow each year and the maximum
extent of land affected by such episodic cultivation. We envisage using
several different offline carbon cycle models for this purpose in order to
take account of uncertainties associated with differences between the
carbon cycle models. The carbon cycle simulations will be driven by climate
outputs (temperature, precipitation, and cloud cover) from an existing
transient climate simulation made with the ECHAM model (Fischer and
Jungclaus, 2011) and <inline-formula><mml:math id="M27" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> prescribed from ice core records. The <inline-formula><mml:math id="M28" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> emission estimates from these two simulations will then be evaluated using C budget constraints. This evaluation will allow us to pinpoint potential discrepancies between known terrestrial C balance changes and estimated LULC <inline-formula><mml:math id="M29" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> emissions in given periods over the Holocene.</p>
</sec>
<sec id="Ch1.S8">
  <label>8</label><title>Implementation of LULC in Earth system model simulations</title>
      <p id="d1e986">We propose a series of simulations to examine the impact of LULC using the
revised LULC scenarios from LandCover6k and building on climate model
experiments that are currently being run either in CMIP6-PMIP4
(<italic>midHolocene</italic>, <italic>past1000</italic>) or within PMIP although not formally included as CMIP6-PMIP4 experiments.</p>

<?xmltex \floatpos{t}?><table-wrap id="Ch1.T1" specific-use="star"><?xmltex \currentcnt{1}?><label>Table 1</label><caption><p id="d1e998">Boundary conditions for CMIP6-PMIP4 and the mid-Holocene LULC experiments. The boundary conditions for the CMIP6-PMIP4 <italic>piControl</italic> and <italic>midHolocene</italic> are described in Otto-Bleisner et al. (2017) and are given here for completeness.</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="5">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="left"/>
     <oasis:colspec colnum="3" colname="col3" align="left"/>
     <oasis:colspec colnum="4" colname="col4" align="left"/>
     <oasis:colspec colnum="5" colname="col5" align="left"/>
     <oasis:thead>
       <oasis:row>
         <oasis:entry namest="col1" nameend="col2" align="center">Boundary conditions </oasis:entry>
         <oasis:entry colname="col3">1850 CE</oasis:entry>
         <oasis:entry colname="col4">6 ka</oasis:entry>
         <oasis:entry colname="col5">6 ka LULC</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3">(DECK</oasis:entry>
         <oasis:entry colname="col4">(<italic>midHolocene</italic>)</oasis:entry>
         <oasis:entry colname="col5">(<italic>midHoloceneLULC</italic>)</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3"><italic>piControl</italic>)</oasis:entry>
         <oasis:entry colname="col4"/>
         <oasis:entry colname="col5"/>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row>
         <oasis:entry colname="col1">Orbital</oasis:entry>
         <oasis:entry colname="col2">Eccentricity</oasis:entry>
         <oasis:entry colname="col3">0.016764</oasis:entry>
         <oasis:entry colname="col4">0.018682</oasis:entry>
         <oasis:entry colname="col5">0.018682</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">parameters</oasis:entry>
         <oasis:entry colname="col2">Obliquity</oasis:entry>
         <oasis:entry colname="col3">23.459</oasis:entry>
         <oasis:entry colname="col4">24.105</oasis:entry>
         <oasis:entry colname="col5">24.105</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2">Perihelion – 180</oasis:entry>
         <oasis:entry colname="col3">100.33</oasis:entry>
         <oasis:entry colname="col4">0.87</oasis:entry>
         <oasis:entry colname="col5">0.87</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2">Vernal equinox</oasis:entry>
         <oasis:entry colname="col3">Noon, 21 March</oasis:entry>
         <oasis:entry colname="col4">Noon, 21 March</oasis:entry>
         <oasis:entry colname="col5">Noon, 21 March</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Greenhouse</oasis:entry>
         <oasis:entry colname="col2">Carbon dioxide (ppm)</oasis:entry>
         <oasis:entry colname="col3">284.3</oasis:entry>
         <oasis:entry colname="col4">264.4</oasis:entry>
         <oasis:entry colname="col5">264.4</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">gases</oasis:entry>
         <oasis:entry colname="col2">Methane (ppb)</oasis:entry>
         <oasis:entry colname="col3">808.2</oasis:entry>
         <oasis:entry colname="col4">597.0</oasis:entry>
         <oasis:entry colname="col5">597.0</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2">Nitrous oxide (ppb)</oasis:entry>
         <oasis:entry colname="col3">273.0</oasis:entry>
         <oasis:entry colname="col4">262.0</oasis:entry>
         <oasis:entry colname="col5">262.0</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2">Other GHGs</oasis:entry>
         <oasis:entry colname="col3">DECK <italic>piControl</italic></oasis:entry>
         <oasis:entry colname="col4">0</oasis:entry>
         <oasis:entry colname="col5">0</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Other</oasis:entry>
         <oasis:entry colname="col2">Solar constant</oasis:entry>
         <oasis:entry colname="col3">TSI: 1360.747</oasis:entry>
         <oasis:entry colname="col4">As <italic>piControl</italic></oasis:entry>
         <oasis:entry colname="col5">As <italic>piControl</italic></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">boundary</oasis:entry>
         <oasis:entry colname="col2">Palaeogeography</oasis:entry>
         <oasis:entry colname="col3">Modern</oasis:entry>
         <oasis:entry colname="col4">As <italic>piControl</italic></oasis:entry>
         <oasis:entry colname="col5">As <italic>piControl</italic></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">conditions</oasis:entry>
         <oasis:entry colname="col2">Ice sheets</oasis:entry>
         <oasis:entry colname="col3">Modern</oasis:entry>
         <oasis:entry colname="col4">As <italic>piControl</italic></oasis:entry>
         <oasis:entry colname="col5">As <italic>piControl</italic></oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2">Vegetation</oasis:entry>
         <oasis:entry colname="col3">Interactive</oasis:entry>
         <oasis:entry colname="col4">Interactive</oasis:entry>
         <oasis:entry colname="col5">Pasture and crop;</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"/>
         <oasis:entry colname="col5">distribution prescribed</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"/>
         <oasis:entry colname="col5">from a revised scenario</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3">DECK <italic>piControl</italic></oasis:entry>
         <oasis:entry colname="col4">As <italic>piControl</italic></oasis:entry>
         <oasis:entry colname="col5">Pasture and crop;</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"/>
         <oasis:entry colname="col5">distribution prescribed</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"/>
         <oasis:entry colname="col5">from a revised scenario</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2">Aerosols</oasis:entry>
         <oasis:entry colname="col3">interactive</oasis:entry>
         <oasis:entry colname="col4">Interactive</oasis:entry>
         <oasis:entry colname="col5">Interactive</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3">DECK <italic>piControl</italic></oasis:entry>
         <oasis:entry colname="col4">As <italic>piControl</italic></oasis:entry>
         <oasis:entry colname="col5">As <italic>piControl</italic></oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

      <p id="d1e1425">The <italic>midHolocene</italic> (and its corresponding <italic>piControl</italic>) simulation is one of the PMIP entry cards in the CMIP6-PMIP4 experiments (Kageyama et al., 2018; Otto-Bliesner et al., 2017), and it is therefore logical to propose this period for LULC simulations. The LULC sensitivity experiment (<?xmltex \hack{\mbox\bgroup}?><italic>midHoloceneLULC</italic><?xmltex \hack{\egroup}?>) should therefore follow the CMIP6-PMIP4 protocol; that is, it should be run with the same climate model components and following the same protocols for implementing external forcings as used in the two CMIP6-PMIP4 experiments (Table 1). Thus, if the <italic>piControl</italic> and <italic>midHolocene</italic> simulations are run with interactive (dynamic) vegetation, then the <italic>midHoloceneLULC</italic> experiment should also be run with dy<?pagebreak page816?>namic vegetation in regions where there is no LULC change. For most models, this means that the LULC forcing is imposed as a fraction of the grid cell and the remaining fraction of the grid cell has simulated natural vegetation. These new mid-Holocene simulations would allow for a better understanding of the relationship between climate changes and land–surface feedbacks (including snow albedo feedbacks), as well as the role of water recycling at a regional scale. Thus, modelling groups who are running the <italic>midHolocene</italic> experiment with a fully interactive carbon cycle could also run the LULC experiment allowing atmospheric <inline-formula><mml:math id="M30" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> to evolve interactively, subject to the simulated ocean and land C balance.</p>
      <p id="d1e1464">The real strength of the revised LULC scenarios is to provide boundary
conditions for transient climate model simulations. The CMIP6-PMIP4 simulation of 850–1850 CE (<italic>past1000</italic>) already incorporates LULC changes as a forcing (Jungclaus et al., 2017) based on a harmonized dataset that provides LULC changes from 850 through 2015 CE (Hurtt et al., 2017), which in turn draws on output from the HYDE3.2 scenario (Klein Goldewijk et al., 2017a). The <italic>past1000</italic> protocol (Jungclaus et al., 2017) acknowledges that this default land use dataset is at the lower end of the spread in estimates of early agricultural area indicated by other LULC scenarios and recommends that modelling groups run additional sensitivity experiments using alternative maximum and minimum scenarios. The revised LULC scenarios created by LandCover6k could be used as an alternative to these maximum and minimum scenarios. Other than the substitution of the LandCover6k scenario, the specifications of other forcings would then follow the recommendations for the CMIP6-PMIP4 <italic>past1000</italic> simulation.</p>
      <p id="d1e1476">A transient climate simulation for a longer period of the Holocene would
provide a more stringent test of the impact of LULC on the coupled Earth
system. We suggest that this transient simulation (<italic>holotrans</italic>) should start from the pre-existing <italic>midHolocene</italic> simulation to capitalize on the fact that the <italic>midHolocene</italic> simulation has been spun up sufficiently long (Otto-Bleisner et al., 2017) to ensure that the ocean and land carbon cycle is in equilibrium at the start of the transient experiment (Table 2). In order to be consistent with the CMIP6-PMIP4 <italic>midHolocene</italic> protocol (Otto-Bleisner et al., 2017), changes in orbital forcing should be specified from Berger and Loutre (1991), and year-by-year changes in <inline-formula><mml:math id="M31" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula>, <inline-formula><mml:math id="M32" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CH</mml:mi><mml:mn mathvariant="normal">4</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula>, and <inline-formula><mml:math id="M33" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">N</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub><mml:mi mathvariant="normal">O</mml:mi></mml:mrow></mml:math></inline-formula> should be specified following Joos and Spahni (2008). LULC changes should be implemented by imposing crop and pasture area through time as specified in the revised LULC scenarios; elsewhere, the simulated vegetation should be active. It will be necessary to run the Holocene transient climate simulation in two steps. A first simulation (<italic>holotrans_LULC</italic>) should be run using prescribed atmospheric <inline-formula><mml:math id="M34" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> concentration, even though the carbon cycle is fully interactive, because this will establish the consistency of the carbon cycle in the land surface model. However, once this is done it will be possible to rerun the simulations with interactive <inline-formula><mml:math id="M35" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> emissions. Table 3 provides a summary of the proposed ESM simulations.</p>

<?xmltex \floatpos{t}?><table-wrap id="Ch1.T2" specific-use="star"><?xmltex \currentcnt{2}?><label>Table 2</label><caption><p id="d1e1555">Boundary conditions for baseline PMIP Holocene transient (6 ka BP to 1850 CE) and LULC transient simulations.</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="5">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="left"/>
     <oasis:colspec colnum="3" colname="col3" align="left"/>
     <oasis:colspec colnum="4" colname="col4" align="left"/>
     <oasis:colspec colnum="5" colname="col5" align="left"/>
     <oasis:thead>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3">Mode</oasis:entry>
         <oasis:entry colname="col4">Source or value</oasis:entry>
         <oasis:entry colname="col5">LULC experiment</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Orbital parameters</oasis:entry>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3">Transient</oasis:entry>
         <oasis:entry colname="col4"/>
         <oasis:entry colname="col5">As baseline simulation</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Greenhouse gases</oasis:entry>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M36" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3">Transient</oasis:entry>
         <oasis:entry colname="col4">Dome C</oasis:entry>
         <oasis:entry colname="col5">As baseline simulation</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M37" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CH</mml:mi><mml:mn mathvariant="normal">4</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4">Combined EPICA &amp; GISP record</oasis:entry>
         <oasis:entry colname="col5">As baseline simulation</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2"><inline-formula><mml:math id="M38" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">N</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub><mml:mi mathvariant="normal">O</mml:mi></mml:mrow></mml:math></inline-formula></oasis:entry>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4">Combined EPICA NGRIP, &amp; TALDICE record</oasis:entry>
         <oasis:entry colname="col5">As baseline simulation</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Solar forcing</oasis:entry>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3">Transient</oasis:entry>
         <oasis:entry colname="col4">Steinhilber et al. (2012)</oasis:entry>
         <oasis:entry colname="col5">As baseline simulation</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Volcanic forcing</oasis:entry>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3">Transient</oasis:entry>
         <oasis:entry colname="col4">To be determined</oasis:entry>
         <oasis:entry colname="col5">As baseline simulation</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Palaeogeography</oasis:entry>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3">Constant at PI values</oasis:entry>
         <oasis:entry colname="col4">Modern</oasis:entry>
         <oasis:entry colname="col5">As baseline simulation</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Ice sheets</oasis:entry>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3">Constant at PI values</oasis:entry>
         <oasis:entry colname="col4">Modern</oasis:entry>
         <oasis:entry colname="col5">As baseline simulation</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Vegetation</oasis:entry>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3">Interactive</oasis:entry>
         <oasis:entry colname="col4"/>
         <oasis:entry colname="col5">LC6k transient pasture and</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3"/>
         <oasis:entry colname="col4"/>
         <oasis:entry colname="col5">crop distribution imposed</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1">Aerosols</oasis:entry>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3">Constant at PI values</oasis:entry>
         <oasis:entry colname="col4"/>
         <oasis:entry colname="col5">As baseline simulation</oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

<?xmltex \floatpos{t}?><table-wrap id="Ch1.T3" specific-use="star"><?xmltex \currentcnt{3}?><label>Table 3</label><caption><p id="d1e1809">Summary of proposed simulations.</p></caption><oasis:table frame="topbot"><oasis:tgroup cols="3">
     <oasis:colspec colnum="1" colname="col1" align="left"/>
     <oasis:colspec colnum="2" colname="col2" align="left"/>
     <oasis:colspec colnum="3" colname="col3" align="left"/>
     <oasis:thead>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1">Name</oasis:entry>
         <oasis:entry colname="col2">Mode</oasis:entry>
         <oasis:entry colname="col3">Purpose</oasis:entry>
       </oasis:row>
     </oasis:thead>
     <oasis:tbody>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1"><italic>piControl</italic></oasis:entry>
         <oasis:entry colname="col2">Equilibrium</oasis:entry>
         <oasis:entry colname="col3">Standard CMIP6-PMIP4 simulation</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1"><italic>midHolocene</italic></oasis:entry>
         <oasis:entry colname="col2">Equilibrium</oasis:entry>
         <oasis:entry colname="col3">Standard CMIP6-PMIP4 simulation</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1"><italic>midHoloceneLULC</italic></oasis:entry>
         <oasis:entry colname="col2">Equilibrium</oasis:entry>
         <oasis:entry colname="col3">Sensitivity to LULC changes</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><italic>holotrans</italic></oasis:entry>
         <oasis:entry colname="col2">Transient</oasis:entry>
         <oasis:entry colname="col3">Baseline fully transient simulation from</oasis:entry>
       </oasis:row>
       <oasis:row rowsep="1">
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3">6 ka onwards, with no LULC</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"><italic>holotrans_LULC</italic></oasis:entry>
         <oasis:entry colname="col2">Transient</oasis:entry>
         <oasis:entry colname="col3">Fully transient simulation from 6 ka</oasis:entry>
       </oasis:row>
       <oasis:row>
         <oasis:entry colname="col1"/>
         <oasis:entry colname="col2"/>
         <oasis:entry colname="col3">onwards, with LULC imposed</oasis:entry>
       </oasis:row>
     </oasis:tbody>
   </oasis:tgroup></oasis:table></table-wrap>

      <?pagebreak page817?><p id="d1e1927"><?xmltex \hack{\newpage}?>Unlike the situation for the mid-Holocene, wherein there is a global climate
benchmark dataset (Bartlein et al., 2011) which provides reconstructions of
multiple bioclimatic variables of seasonal temperature and moisture, the
opportunities for quantitative evaluation of the <italic>holotrans</italic> simulated climate are more limited. Seasonal temperature reconstructions are available for Europe (Davis et al., 2003) and North America (Viau et al., 2006; Viau and Gajewski, 2009). Although there is a new global dataset that provides global temperature reconstructions for the Holocene (Kaufman et al., 2020), it is based on only 472 terrestrial records worldwide, and the results for zonally averaged temperature changes are therefore likely to be more robust than the regional details. There are also time series reconstructions for individual sites outside these two regions (e.g. Nakagawa et al., 2002;
Wilmshurst et al., 2007; Ortega-Rosas et al., 2008). Furthermore, the simulated time course of <inline-formula><mml:math id="M39" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> emissions can be compared to the ice core records.</p>
      <p id="d1e1946">The CMIP6-PMIP4 <italic>midHolocene</italic> simulations are stylized experiments lacking several potential forcings (in addition to LULC), including changes in atmospheric dust loading, solar irradiance, and volcanic forcing. We suggest that additional sensitivity tests could be run to take these additional forcings into account. In the case of solar and volcanic forcing, this would also ensure that the transient <italic>holotrans</italic> simulations mesh seamlessly with the <italic>past1000</italic> simulation. Changes in solar variability during the Holocene should be specified from Steinhilber et al. (2012). There are records of volcanic forcing for the past 2000 years (Sigl et al., 2015; Toohey and Sigl, 2017), and these are used in the <italic>past1000</italic> simulation. Observationally constrained estimates of
volcanic stratospheric aerosol for the Holocene are currently under development (Michael Sigl, personal communication, 2019) and could be implemented as an additional sensitivity experiment when available. Changes in atmospheric dust loading are not included in the <italic>past1000</italic> simulation but are important during the earlier part of the Holocene (Pausata et al., 2016; Tierney et al., 2017; Messori et al., 2019). Although continuous reconstructions of dust loading through the Holocene are not available, it would be possible to use estimates for particular time slices (Egerer et al., 2018) to test the sensitivity to this forcing.</p>
</sec>
<?pagebreak page818?><sec id="Ch1.S9" sec-type="conclusions">
  <label>9</label><title>Outcomes and perspectives</title>
      <p id="d1e1972">LandCover6k has developed a scheme for using archaeological information to
improve existing scenarios of LULC changes during the Holocene, specifically by using archaeological data to provide better estimates of regional population changes through time, better information on the date of the initiation of agriculture in a region, more regionally specific information about the type of land use, and more nuanced information about land use per capita than currently implemented in the LULC scenarios generated by HYDE and KK10. While the final global datasets are still in production, fast-track priority products have been created and their impact on current scenarios is being tested.</p>
      <p id="d1e1975">Although the work of LandCover6k will provide more solid knowledge about
anthropogenic modification of the landscape, some information will inevitably be missing and some key regions will be poorly covered. There will still be large uncertainties associated with revised LULC scenarios, even though these will be based on more solid evidence than the existing LULC scenarios. Documenting the uncertainties in the archaeological inputs and their impacts on the revised scenarios is an important goal of the LandCover6k project. We propose using the information about the uncertainties in the archaeological data sources to generate multiple LULC scenarios comparable to the “low-end” and “high-end” scenarios used for e.g. future projections. Furthermore, we have proposed a series of tests that will help to evaluate the realism of the final scenarios based on independent evidence from pollen-based reconstructions of land cover, reconstructions of climate, and carbon cycle constraints. These tests should help in identifying which of the potential LULC reconstructions are most realistic and in constraining the sources of uncertainty.</p>
      <p id="d1e1978">We have proposed the use of offline carbon cycle simulations solely as a test of the realism of the revised LULC scenarios. Quantifying the LULC contribution to <inline-formula><mml:math id="M40" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> emissions during the Holocene would require
additional simulations in which other forcings (climate, atmospheric
<inline-formula><mml:math id="M41" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula>, insolation) are kept constant. The difference in simulated total terrestrial C storage between these simulations and LULC simulations
provides an estimate of <italic>primary emissions</italic> (Pongratz et al., 2014) and avoids additional model uncertainty regarding the sensitivity of land C storage to atmospheric <inline-formula><mml:math id="M42" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> or climate being included in emission estimates. There are other sensitivity tests that would be useful. For example, vegetation–carbon cycle models differ in their ability to account for gross land use transitions within grid cells (Arneth et al., 2017). This is critical for simulating the effects of non-permanent agriculture, whereby land is simultaneously abandoned and reclaimed within the extent of a model grid cell. Such shifting cultivation-type agriculture implies forest degradation in areas recovering from previous land use and leads to substantially higher LULC emissions compared to model estimates wherein only net land use changes are accounted for (Shevliakova et al., 2009). It would therefore be interesting to run additional simulations accounting for net land use change and indeed separating out the effects of wood harvesting and shifting cultivation.</p>
      <p id="d1e2017">We anticipate that it will be possible to incorporate realistic LULC scenarios for the mid-Holocene as part of the climate model sensitivity experiments planned during PMIP4. Such experiments will complement the
CMIP6-PMIP4 baseline model experiments by providing insights into whether
discrepancies between simulated and observed 6 ka climate could be the result of incorrect specification of the land–surface boundary conditions.
However, the incorporation of archaeological information into LULC scenarios
clearly makes it possible to target other interesting periods for such
experiments, for example to explore whether land use changes played a role in
abrupt events such as the 4.2 ka event, the impact of population declines in the Americas as a consequence of European colonization (1500–1750 CE), or the changes in land use globally during the industrial era (post-1850 CE).</p>
      <p id="d1e2021">In addition to providing a protocol for the PMIP 6 ka sensitivity experiments, we have devised a protocol for implementing the optimal LULC
reconstructions for the Holocene in transient climate model or ESM experiments. The goal here is to provide one of the necessary forcings that
could be used for transient simulations in future phases of PMIP. This will
allow for an assessment of LULC in these simulations and therefore help address
issues that are a focus for other MIPs e.g. LUMIP or LS3MIP. When these new
forcings are created, they will be made available through the PMIP4 website
(<uri>https://pmip4.lsce.ipsl.fr/doku.php/exp_design:lgm</uri>, last access: 14 February 2020, PMIP4 repository, 2017) and the Earth System Grid Federation (ESGF) Input4MIPS repository (<uri>https://esgf-node.llnl.gov/projects/input4mips/</uri>, last access: 14 February 2020, with details provided in the “input4MIPs summary” link). Modelling groups who run either equilibrium or transient climate model experiments following this protocol are encouraged to follow the standard CMIP protocol for archiving their simulations through the ESGF.</p>
</sec>

      
      </body>
    <back><notes notes-type="codedataavailability"><title>Code and data availability</title>

      <p id="d1e2034">The data used for Fig. 1 are publicly available. The HYDE3.2 data can be downloaded from: <ext-link xlink:href="https://doi.org/10.17026/dans-25g-gez3" ext-link-type="DOI">10.17026/dans-25g-gez3</ext-link> (Klein Goldewijk, 2017). The KK10 data can be downloaded from: <ext-link xlink:href="https://doi.org/10.1594/PANGAEA.871369" ext-link-type="DOI">10.1594/PANGAEA.871369</ext-link> (Kaplan and Krumhardt, 2011). The code and data used to generate Fig. 1 are available from: <uri>https://github.com/jedokaplan/ALCC_comparison_figure</uri> (Kaplan, 2020). The data and code used to generate Fig. 3 are available from: <uri>https://github.com/mavdlind/GMD</uri> (Vander Linden, 2020). The data and code used  to generate Fig. 5 are available from: <ext-link xlink:href="https://doi.org/10.5281/zenodo.3625226" ext-link-type="DOI">10.5281/zenodo.3625226</ext-link> (Lewin and Whitehouse, 2020. The European pollen-based reconstructions used in Fig. 7 are available at:
<ext-link xlink:href="https://doi.org/10.1594/PANGAEA.897303" ext-link-type="DOI">10.1594/PANGAEA.897303</ext-link> (Gaillard, 2019). The pollen data used to generate the Siberian reconstructions are available from: <ext-link xlink:href="https://doi.org/10.1594/PANGAEA.898616" ext-link-type="DOI">10.1594/PANGAEA.898616</ext-link>.<?pagebreak page819?> An earlier version of this figure was published in Dawson et al. (2018). The code used to generate Fig. 7 is available from: <ext-link xlink:href="https://doi.org/10.5281/zenodo.3604328" ext-link-type="DOI">10.5281/zenodo.3604328</ext-link> (Dawson, 2019). The
pollen-based reconstructions used in the generation of Fig. 8 are available from: <ext-link xlink:href="https://doi.org/10.5281/zenodo.3601028" ext-link-type="DOI">10.5281/zenodo.3601028</ext-link> (Harrison, 2020a). The climate model outputs used to generate Fig. 8 are available from: <ext-link xlink:href="https://doi.org/10.5281/zenodo.3601040" ext-link-type="DOI">10.5281/zenodo.3601040</ext-link> (Harrison, 2020b). The code used to generate Fig. 8 is available from: <ext-link xlink:href="https://doi.org/10.5281/zenodo.3601011" ext-link-type="DOI">10.5281/zenodo.3601011</ext-link> (Harrison, 2020c).</p>
  </notes><notes notes-type="authorcontribution"><title>Author contributions</title>

      <p id="d1e2075">SPH, MJG, BDS, MVL, and KKG wrote the first draft. SPH, PB, and FSRP contributed to the design of the climate model experiments; BS and TK contributed to the design of the carbon cycle simulations. The figures were contributed by JK (Fig. 1), BS (Figs. 2 and 9), MVL (Fig. 3), OB (Fig. 4), NJW (Fig. 5), KKG (Fig. 6), AD (Fig. 7), and SPH (Fig. 8). All authors contributed to the final version of the paper.</p>
  </notes><notes notes-type="competinginterests"><title>Competing interests</title>

      <p id="d1e2081">The authors declare that they have no conflict of interest.</p>
  </notes><notes notes-type="sistatement"><title>Special issue statement</title>

      <p id="d1e2087">This article is part of the special issue “Paleoclimate Modelling Intercomparison Project phase 4 (PMIP4) (CP/GMD inter-journal SI)”. It is not associated with a conference.</p>
  </notes><ack><title>Acknowledgements</title><p id="d1e2093">LandCover6k is a working group of the Past Global Changes (PAGES) programme, which in turn received support from the Swiss Academy of Sciences. We thank PAGES for their support for this activity. The land use group also received funding under the Holocene Global Landuse International Focus Group of INQUA. SPH acknowledges funding from the European Research Council for “GC2.0: Unlocking the past for a clearer future”. Marie-José Gaillard thanks the Swedish Strategic Research Area MERGE (Modelling the Regional and Global Earth System) and Linnaeus University's faculty of Health and Life Sciences (Kalmar, Sweden) for financial support. We thank Joy Singarayer for providing the climate model outputs that were used to generate Fig. 8 and Guangqi Li for assistance in producing this figure. BDS was funded by ERC H2020-MSCA-IF-2015. The dataset for Fig. 5 was generated from the “Cultivating Societies: Assessing the Evidence for Agriculture in Neolithic Ireland” project, supported by the Heritage Council, Ireland, under the INSTAR programme 2008–2010.</p></ack><notes notes-type="financialsupport"><title>Financial support</title>

      <p id="d1e2098">This research has been supported by the European Research Council (grant nos. 694481 and 701329) and the Heritage Council, Ireland (grant no. 16682).</p>
  </notes><notes notes-type="reviewstatement"><title>Review statement</title>

      <p id="d1e2104">This paper was edited by Richard Neale and reviewed by two anonymous referees.</p>
  </notes><ref-list>
    <title>References</title>

      <ref id="bib1.bib1"><label>1</label><?label 1?><mixed-citation>Arneth, A., Sitch, S., Pongratz, J., Stocker, B. D., Ciais, P., Poulter, B.,
Bayer, A. D., Bondeau, A., Calle, L., Chini, L. P., Gasser, T., Fader, M.,
Friedlingstein, P., Kato, E., Li, W., Lindeskog, M., Nabel, J. E. M. S., Pugh, T. A. M., Robertson, E., Viovy, N., Yue, C., and Zaehle, S.: Historical carbon dioxide emissions caused by land-use changes are possibly larger than
assumed, Nat. Geosci., 10, 79–84, <ext-link xlink:href="https://doi.org/10.1038/ngeo2882" ext-link-type="DOI">10.1038/ngeo2882</ext-link>, 2017.</mixed-citation></ref>
      <ref id="bib1.bib2"><label>2</label><?label 1?><mixed-citation>
Arneth, A., Denton, F., Agus, F., Elbehri, A., Erb, K., Elasha, B. O., Rahimi, M., Rounsevell, M., Spence, A., and Valentini, R.: IPCC Special
Report on Climate Change, Desertification, Land Degradation, Sustainable
Land Management, Food Security, and Greenhouse gas fluxes in Terrestrial
Ecosystems, 2019.</mixed-citation></ref>
      <ref id="bib1.bib3"><label>3</label><?label 1?><mixed-citation>Balsera, V., Díaz-del-Río, P., Gilman, A., Uriarte, A., and Vicent, J. M.: Approaching the demography of late prehistoric Iberia through summed calibrated probability distributions (7000–2000 cal BC), Quatern. Int.,
208–211, <ext-link xlink:href="https://doi.org/10.1016/j.quaint.2015.06.022" ext-link-type="DOI">10.1016/j.quaint.2015.06.022</ext-link>, 2015.</mixed-citation></ref>
      <ref id="bib1.bib4"><label>4</label><?label 1?><mixed-citation>Bartlein, P. J., Harrison, S. P., Brewer, S., Connor, S., Davis, B. A. S.,
Gajewski, K., Guiot, J., Harrison-Prentice, T. I., Henderson, A., Peyron, O., Prentice, I. C., Scholze, M., Seppä, H., Shuman, B., Sugita, S., Thompson, R. S., Viau, A., Williams, J., and Wu, H.: Pollen-based continental climate reconstructions at 6 and 21 ka: a global synthesis, Clim. Dynam., 37, 775–802, <ext-link xlink:href="https://doi.org/10.1007/s00382-010-0904-1" ext-link-type="DOI">10.1007/s00382-010-0904-1</ext-link>, 2011.</mixed-citation></ref>
      <ref id="bib1.bib5"><label>5</label><?label 1?><mixed-citation>Bartlein, P. J., Harrison, S. P., and Izumi, K.: Underlying causes of Eurasian mid-continental aridity in simulations of mid-Holocene climate, Geophys. Res. Lett., 44, 9020–9028, <ext-link xlink:href="https://doi.org/10.1002/2017GL074476" ext-link-type="DOI">10.1002/2017GL074476</ext-link>, 2017.</mixed-citation></ref>
      <ref id="bib1.bib6"><label>6</label><?label 1?><mixed-citation>Barton, C. M., Ullah, I. I., and Bergin, S.: Land use, water and Mediterranean landscapes: modelling long-term dynamics of complex socio-ecological systems, Philos. T. Roy. Soc. A,368, 5275–5297, <ext-link xlink:href="https://doi.org/10.1098/rsta.2010.0193" ext-link-type="DOI">10.1098/rsta.2010.0193</ext-link>, 2010.</mixed-citation></ref>
      <ref id="bib1.bib7"><label>7</label><?label 1?><mixed-citation>Berger, A.  and Loutre, M.-F.: Insolation values for the climate of the last
10 million of years, Quaternary Sci. Rev., 10, 297–317, <ext-link xlink:href="https://doi.org/10.1016/0277-3791(91)90033-Q" ext-link-type="DOI">10.1016/0277-3791(91)90033-Q</ext-link>, 1991.</mixed-citation></ref>
      <ref id="bib1.bib8"><label>8</label><?label 1?><mixed-citation>
Bishop, R. R., Church, M. J., and Rowley-Conwy, P. A.: Firewood, food and human niche construction: the potential role of Mesolithic hunter–gatherers in actively structuring Scotland's woodlands, Quaternary Sci. Rev., 108, 51–75, 2015.</mixed-citation></ref>
      <ref id="bib1.bib9"><label>9</label><?label 1?><mixed-citation>Bocquet-Appel, J.-P., Naji, S., Vander Linden, M., and Kozłowsi, J. K.:
Detection of diffusion and contact zones of early farming in Europe from the
space-time distribution of <inline-formula><mml:math id="M43" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi/><mml:mn mathvariant="normal">14</mml:mn></mml:msup><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula> dates, J. Arch. Sci., 36, 807–820,
<ext-link xlink:href="https://doi.org/10.1016/j.jas.2008.11.004" ext-link-type="DOI">10.1016/j.jas.2008.11.004</ext-link>, 2009.</mixed-citation></ref>
      <ref id="bib1.bib10"><label>10</label><?label 1?><mixed-citation>Crema, E. R., Habu, J., Kobayashi, K., and Madella, M.: Summed probability
distribution of <inline-formula><mml:math id="M44" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi/><mml:mn mathvariant="normal">14</mml:mn></mml:msup><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula> dates suggests regional divergences in the
population dynamics of the Jomon period in eastern Japan, PlosOne, 11,
e0154809, <ext-link xlink:href="https://doi.org/10.1371/journal.pone.0154809" ext-link-type="DOI">10.1371/journal.pone.0154809</ext-link>, 2016.</mixed-citation></ref>
      <ref id="bib1.bib11"><label>11</label><?label 1?><mixed-citation>
Davis, B. A. S., Brewer, S., Stevenson, A. C., Guiot, J., and Juggins, S.: The temperature of Europe during the Holocene reconstructed from pollen data,
Quaternary Sci. Rev., 22, 1701–1716, 2003.</mixed-citation></ref>
      <ref id="bib1.bib12"><label>12</label><?label 1?><mixed-citation>Davison, K., Dolukhanov, P., Sarson, G. R., and Shukurov, P.: The role of
waterways in the spread of the Neolithic, J. Arch. Sci., 33, 641–652, <ext-link xlink:href="https://doi.org/10.1016/j.jas.2005.09.017" ext-link-type="DOI">10.1016/j.jas.2005.09.017</ext-link>, 2006.</mixed-citation></ref>
      <ref id="bib1.bib13"><label>13</label><?label 1?><mixed-citation>Dawson, A.: Code to reproduce Figure 7 from Harrison et al., GMD, <ext-link xlink:href="https://doi.org/10.5281/zenodo.3604328" ext-link-type="DOI">10.5281/zenodo.3604328</ext-link>, 2019.</mixed-citation></ref>
      <?pagebreak page820?><ref id="bib1.bib14"><label>14</label><?label 1?><mixed-citation>Dawson, A., Paciorek, C. J., McLachlan, J. S., Goring, S., Williams, J. W., and Jackson, S. T.: Quantifying pollen-vegetation relationships to reconstruct ancient forests using 19th-century forest composition and pollen data, Quaternary Sci. Rev., 137, 156–175, <ext-link xlink:href="https://doi.org/10.1016/j.quascirev.2016.01.012" ext-link-type="DOI">10.1016/j.quascirev.2016.01.012</ext-link>, 2016.</mixed-citation></ref>
      <ref id="bib1.bib15"><label>15</label><?label 1?><mixed-citation>Dawson, A., Cao, X., Chaput, M., Hopla, E., Li, F., Edwards, M., Fyfe, R.,
Gajewski, K., Goring, S. J., Herzschuh, U., Mazier, F., Sugita, S., Williams,
J. W., Xu, Q., and Gaillard, M.-J.: Finding the magnitude of human induced
Northern Hemisphere land-cover transformation between 6 and 0.2 ka BP, PAGES Mag., 26, 34–35, <ext-link xlink:href="https://doi.org/10.22498/pages.26.1.34" ext-link-type="DOI">10.22498/pages.26.1.34</ext-link>, 2018.</mixed-citation></ref>
      <ref id="bib1.bib16"><label>16</label><?label 1?><mixed-citation>Egerer, S., Claussen, M., and Reick, C.: Rapid increase in simulated North
Atlantic dust deposition due to fast change of northwest African landscape
during the Holocene, Clim. Past, 14, 1051–1066,
<ext-link xlink:href="https://doi.org/10.5194/cp-14-1051-2018" ext-link-type="DOI">10.5194/cp-14-1051-2018</ext-link>, 2018.</mixed-citation></ref>
      <ref id="bib1.bib17"><label>17</label><?label 1?><mixed-citation>
Elsig, J., Schmitt, J., Leuenberger, D., Schneider, R., Eyer, M.,
Leuenberger, M., Joos, F., Fischer, H., and Stocker, T. F.: Stable isotope
constraints on Holocene carbon cycle changes from an Antarctic ice core,
Nature, 461, 507–510, 2009.</mixed-citation></ref>
      <ref id="bib1.bib18"><label>18</label><?label 1?><mixed-citation>Eyring, V., Bony, S., Meehl, G. A., Senior, C. A., Stevens, B., Stouffer, R.
J., and Taylor, K. E.: Overview of the Coupled Model Intercomparison Project
Phase 6 (CMIP6) experimental design and organization, Geosci. Model Dev., 9,
1937–1958, <ext-link xlink:href="https://doi.org/10.5194/gmd-9-1937-2016" ext-link-type="DOI">10.5194/gmd-9-1937-2016</ext-link>, 2016.</mixed-citation></ref>
      <ref id="bib1.bib19"><label>19</label><?label 1?><mixed-citation>Fischer, N. and Jungclaus, J. H.: Evolution of the seasonal temperature
cycle in a transient Holocene simulation: orbital forcing and sea-ice, Clim.
Past, 7, 1139–1148, <ext-link xlink:href="https://doi.org/10.5194/cp-7-1139-2011" ext-link-type="DOI">10.5194/cp-7-1139-2011</ext-link>, 2011.</mixed-citation></ref>
      <ref id="bib1.bib20"><label>20</label><?label 1?><mixed-citation>Ford, A. and Clarke, K. C.: Linking the past and present of the ancient Maya: lowland land use, population distribution, and density in the Late Classic Period, in: The Oxford Handbook of Historical Ecology and Applied Archaeology, edited by: Isendahl, C. and Stump, D., Oxford University Press, Oxford, <ext-link xlink:href="https://doi.org/10.1093/oxfordhb/9780199672691.013.33" ext-link-type="DOI">10.1093/oxfordhb/9780199672691.013.33</ext-link>, 2015.</mixed-citation></ref>
      <ref id="bib1.bib21"><label>21</label><?label 1?><mixed-citation>Freeman, J., Baggio, J. A., Robinson, E., Byers, D. A., Gayo, E., Finley, J. B., Meyer, J. A., Kelly, R. L., and Anderies, J. M.: Synchronisation of energy consumption by human societies throughout the Holocene, P. Natl. Acad. Sci. USA, 115, 9962–9967, <ext-link xlink:href="https://doi.org/10.1073/pnas.1802859115" ext-link-type="DOI">10.1073/pnas.1802859115</ext-link>, 2018.</mixed-citation></ref>
      <ref id="bib1.bib22"><label>22</label><?label 1?><mixed-citation>Fyfe, R. M., Woodbridge, J. E., and Roberts, N.: From forest to farmland:
pollen-inferred land cover change across Europe using the pseudobiomization
approach, Global Change Biol., 21: 1197–1212, <ext-link xlink:href="https://doi.org/10.1111/gcb.12776" ext-link-type="DOI">10.1111/gcb.12776</ext-link>, 2014.</mixed-citation></ref>
      <ref id="bib1.bib23"><label>23</label><?label 1?><mixed-citation>Gaillard, M.-J.: First pollen-based REVEALS reconstruction of vegetation abundance in Europe for five time windows between 6 k years BP and present, PANGAEA, <ext-link xlink:href="https://doi.org/10.1594/PANGAEA.897303" ext-link-type="DOI">10.1594/PANGAEA.897303</ext-link>, 2019.</mixed-citation></ref>
      <ref id="bib1.bib24"><label>24</label><?label 1?><mixed-citation>Gaillard, M.-J., Sugita, S., Mazier, F., Kaplan, J. O., Trondman, A.-K., Brostroem, A., Hickler, T., Kjellstroem, E., Kunes, P., Lemmen, C., Olofsson, J., Smith, B., and Strandberg, G.: Holocene land-cover reconstructions for studies on land-cover feedbacks, Clim. Past, 6, 483–499,
<ext-link xlink:href="https://doi.org/10.5194/cp-6-483-2010" ext-link-type="DOI">10.5194/cp-6-483-2010</ext-link>, 2010.</mixed-citation></ref>
      <ref id="bib1.bib25"><label>25</label><?label 1?><mixed-citation>Gaillard, M-J., Whitehouse, N., Madella, M., Morrison, K., and von Gunten, L.: Past land use and land cover, PAGES Mag.,26, 1–44,
<ext-link xlink:href="https://doi.org/10.22498/pages.26.1" ext-link-type="DOI">10.22498/pages.26.1</ext-link>, 2018.</mixed-citation></ref>
      <ref id="bib1.bib26"><label>26</label><?label 1?><mixed-citation>Harrison, S. P.: Pollen-based reconstructions of bioclimatic variables at 6000 yr BP, <ext-link xlink:href="https://doi.org/10.5281/zenodo.3601028" ext-link-type="DOI">10.5281/zenodo.3601028</ext-link>, 2020a.</mixed-citation></ref>
      <ref id="bib1.bib27"><label>27</label><?label 1?><mixed-citation>Harrison, S. P.: Climate simulations of impact of LULC on mid-Holocene climate, <ext-link xlink:href="https://doi.org/10.5281/zenodo.3601040" ext-link-type="DOI">10.5281/zenodo.3601040</ext-link>, 2020b.</mixed-citation></ref>
      <ref id="bib1.bib28"><label>28</label><?label 1?><mixed-citation>Harrison, S. P.: Code to compare model outputs and pollen-based reconstructions, <ext-link xlink:href="https://doi.org/10.5281/zenodo.3601011" ext-link-type="DOI">10.5281/zenodo.3601011</ext-link>, 2020c.</mixed-citation></ref>
      <ref id="bib1.bib29"><label>29</label><?label 1?><mixed-citation>
Harrison, S. P., Bartlein, P. J., Izumi, K., Li, G., Annan, J., Hargreaves,
J., Braconnot, P. B., and Kageyama, M.: Evaluation of CMIP5 palaeo-simulations to improve climate projections, Nat. Clim. Change, 5,
735–743, 2015.</mixed-citation></ref>
      <ref id="bib1.bib30"><label>30</label><?label 1?><mixed-citation>
He, F., Vavrus, S. J., Kutzbach, J. E., Ruddiman, W. F., Kaplan, J. O., and
Krumhardt, K. M.: Simulating global and local surface temperature changes due
to Holocene anthropogenic land cover change, Geophys. Res. Lett., 41, 623–631, 2014.</mixed-citation></ref>
      <ref id="bib1.bib31"><label>31</label><?label 1?><mixed-citation>
Hellman, S., Gaillard, M.-J., Broström, A., and Sugita, S.: The REVEALS
model, new tool to estimate past regional plant abundance from pollen data
in large lakes: validation in southern Sweden, J. Quatenary Sci., 22, 1–22,
2008a.</mixed-citation></ref>
      <ref id="bib1.bib32"><label>32</label><?label 1?><mixed-citation>
Hellman, S., Gaillard, M.-J., Broström, A., and Sugita, S.: Effects of
the sampling design and selection of parameter values on pollen-based
quantitative reconstructions of regional vegetation: a case study in southern Sweden using the REVEALS model, Veg. Hist. Archaeobot., 17, 445–460, 2008b.</mixed-citation></ref>
      <ref id="bib1.bib33"><label>33</label><?label 1?><mixed-citation>
Herrero, M., Havlík, P., Valin, H., Notenbaert, A., Rufino, M. C.,
Thornton, P. K., Blümmel, M., Weiss, F., Grace, D., and Obersteiner, M.:
Biomass use, production, feed efficiencies, and greenhouse gas emissions from global livestock systems, P. Natl. Acad. Sci. USA, 110, 20888–20893, 2013.</mixed-citation></ref>
      <ref id="bib1.bib34"><label>34</label><?label 1?><mixed-citation>Hughes, R. E., Weiberg, E., Bonnier, A., Finne, M., and Kaplan, J. O.:
Quantifying land use in past societies from cultural practice and archaeological data, Land, 7, 9, <ext-link xlink:href="https://doi.org/10.3390/land7010009" ext-link-type="DOI">10.3390/land7010009</ext-link>, 2018.</mixed-citation></ref>
      <ref id="bib1.bib35"><label>35</label><?label 1?><mixed-citation>Hurtt, G., Chini, L., Sahajpal, R., Frolking, S., Bodirsky, B. L., Calvin, K., Doelman, J., Fisk, J., Fujimori, S., Klein Goldewijk, K., Hasegawa, T., Havlik, P., Heinimann, A., Humpenöder, F., Jungclaus, J., Kaplan, J., Krisztin, T., Lawrence, D., Lawrence, P., Mertz, O., Pongratz, J., Popp, A., Riahi, K., Shevliakova, E., Stehfest, E., Thornton, P., van Vuuren, D., and Zhang, X.: Harmonization of global land use scenarios (LUH2): SSP585 v2.1f 2015–2100, Earth System Grid Federation, <ext-link xlink:href="https://doi.org/10.22033/ESGF/input4MIPs.1662" ext-link-type="DOI">10.22033/ESGF/input4MIPs.1662</ext-link>, 2017</mixed-citation></ref>
      <ref id="bib1.bib36"><label>36</label><?label 1?><mixed-citation>
Janssen, E., Poblome, J., Claeys, J., Kint, V., Degryse, P., Marinova, E., and Muys, B.: Fuel for debating ancient economies. Calculating wood consumption at urban scale in Roman Imperial times, J. Arch. Sci.: Rep., 11, 592–599, 2017.</mixed-citation></ref>
      <ref id="bib1.bib37"><label>37</label><?label 1?><mixed-citation>
Joos, F. and Spahni, R.: Rates of change in natural and anthropogenic radiative forcing over the past 20,000 years, P. Natl. Acad. Sci. USA, 105,
1425–1430, 2008.</mixed-citation></ref>
      <ref id="bib1.bib38"><label>38</label><?label 1?><mixed-citation>Joos, F., Gerber, S., Prentice, I. C., Otto-Bliesner, B. L., and Valdes, P. J.: Transient simulations of Holocene atmospheric carbon dioxide and terrestrial carbon since the last glacial maximum, Global Biogeochem. Cy., 18, GB2002, <ext-link xlink:href="https://doi.org/10.1029/2003GB002156" ext-link-type="DOI">10.1029/2003GB002156</ext-link>, 2004.</mixed-citation></ref>
      <ref id="bib1.bib39"><label>39</label><?label 1?><mixed-citation>Jungclaus, J. H., Bard, E., Baroni, M., Braconnot, P., Cao, J., Chini, L. P.,
Egorova, T., Evans, M., González-Rouco, J. F., Goosse, H., Hurtt, G. C.,
Joos, F., Kaplan, J. O., Khodri, M., Klein Goldewijk, K., Krivova, N.,
LeGrande, A. N., Lorenz, S. J., Luterbacher, J., Man, W., Maycock, A. C.,
Meinshausen, M., Moberg, A., Muscheler, R., Nehrbass-Ahles, C.,
Otto-Bliesner, B. I., Phipps, S. J., Pongratz, J., Rozanov, E., Schmidt, G. A., Schmidt, H.<?pagebreak page821?>, Schmutz, W., Schurer, A., Shapiro, A. I., Sigl, M., Smerdon,
J. E., Solanki, S. K., Timmreck, C., Toohey, M., Usoskin, I. G., Wagner, S.,
Wu, C.-J., Yeo, K. L., Zanchettin, D., Zhang, Q., and Zorita, E.: The PMIP4
contribution to CMIP6 – Part 3: The last millennium, scientific objective,
and experimental design for the PMIP4 past1000 simulations, Geosci. Model
Dev., 10, 4005–4033, <ext-link xlink:href="https://doi.org/10.5194/gmd-10-4005-2017" ext-link-type="DOI">10.5194/gmd-10-4005-2017</ext-link>, 2017.</mixed-citation></ref>
      <ref id="bib1.bib40"><label>40</label><?label 1?><mixed-citation>Kageyama, M., Braconnot, P., Harrison, S. P., Haywood, A., Jungclaus, J.,
Otto-Bliesner, B., Peterschmitt, J.-Y., Abe-Ouchi, A., Albani, S., Bartlein,
P., Brierley, C., Crucifix, M., Dolan, A., Fernandez-Donado, L., Fischer, H., Hopcroft, P., Ivanovic, R., Lambert, F., Lunt, D., Mahowald, N., Peltier, W. R., Phipps, S., Roche, D., Schmidt, G., Tarasov, L., Valdes, P., Zhang, Q., and Zhou, T.: The PMIP4 contribution to CMIP6 – Part 1: Overview and over-arching analysis plan, Geosci. Model Dev., 11, 1033–1057,
<ext-link xlink:href="https://doi.org/10.5194/gmd-11-1033-2018" ext-link-type="DOI">10.5194/gmd-11-1033-2018</ext-link>, 2018.</mixed-citation></ref>
      <ref id="bib1.bib41"><label>41</label><?label 1?><mixed-citation>Kaplan, J. O.: ALCC_Comparison_Figure, available at: <uri>https://github.com/jedokaplan/ALCC_comparison_figure</uri>, last access: 14 February 2020.</mixed-citation></ref>
      <ref id="bib1.bib42"><label>42</label><?label 1?><mixed-citation>Kaplan, J. O. and Krumhardt, K. M.: The KK10 Anthropogenic Land Cover Change scenario for the preindustrial Holocene, PANGAEA, <ext-link xlink:href="https://doi.org/10.1594/PANGAEA.871369" ext-link-type="DOI">10.1594/PANGAEA.871369</ext-link>, 2011.</mixed-citation></ref>
      <ref id="bib1.bib43"><label>43</label><?label 1?><mixed-citation>
Kaplan, J. O., Krumhardt, K. M., Ellis, E. C., Ruddiman, W. F., Lemmen, C., and Klein Goldewijk, K.: Holocene carbon emissions as a result of anthropogenic land cover change, Holocene, 21, 775–791, 2011.</mixed-citation></ref>
      <ref id="bib1.bib44"><label>44</label><?label 1?><mixed-citation>Kaplan, J. O., Krumhardt, K. M., Gaillard, M.-J., Sugita, S., Trondman, A.-K., Fyfe, R., Marquer, L., Mazier, F., and Nielsen, A. B.: Constraining the deforestation history of Europe: Evaluation of historical land use scenarios with pollen-based land cover reconstructions, Land, 6, 9, <ext-link xlink:href="https://doi.org/10.3390/land6040091" ext-link-type="DOI">10.3390/land6040091</ext-link>, 2017.</mixed-citation></ref>
      <ref id="bib1.bib45"><label>45</label><?label 1?><mixed-citation>
Kaufman, D., McKay, N., Routson, C., Erb, M., Davis, B., Heiri, O., Jaccard,
S., Tierney, J., Dätwyler, C., et al.: A global database of Holocene paleo-temperature records, Scient. Data, in press, 2020.</mixed-citation></ref>
      <ref id="bib1.bib46"><label>46</label><?label 1?><mixed-citation>Kay, A. U., Fuller, D. Q., Neumann, K., Eichhorn, B., Höhn, A., Morin-Rivat, J., Champion, L., Linseele, V., Huysecom, E., Ozainne, S., Lespez, L., Biagetti, S., Madella, M., Salzmann, U., and Kaplan, J. O.:
Diversification, intensification, and specialization: Changing land use in
western Africa from 1800 BC to AD 1500, J. World Prehist., 32, 179–228,
<ext-link xlink:href="https://doi.org/10.1007/s10963-019-09131-2" ext-link-type="DOI">10.1007/s10963-019-09131-2</ext-link>, 2019.</mixed-citation></ref>
      <ref id="bib1.bib47"><label>47</label><?label 1?><mixed-citation>Klein Goldewijk, C. G. M.: Anthropogenic land-use estimates for the Holocene; HYDE 3.2. DANS, <ext-link xlink:href="https://doi.org/10.17026/dans-25g-gez3" ext-link-type="DOI">10.17026/dans-25g-gez3</ext-link>, 2017.</mixed-citation></ref>
      <ref id="bib1.bib48"><label>48</label><?label 1?><mixed-citation>
Klein Goldewijk, K., Beusen, A., van Drecht, G., and de Vos, M.: The HYDE 3.1 spatially explicit database of human induced land use change over the past 12,000 years, Global Ecol. Biogeogr., 20, 73–86, 2011.</mixed-citation></ref>
      <ref id="bib1.bib49"><label>49</label><?label 1?><mixed-citation>Klein Goldewijk, K., Beusen, A., Doelman, J., and Stehfest, E.: Anthropogenic land-use estimates for the Holocene; HYDE 3.2, Earth Syst. Sci. Data, 9, 927–953, <ext-link xlink:href="https://doi.org/10.5194/essd-9-927-2017" ext-link-type="DOI">10.5194/essd-9-927-2017</ext-link>, 2017a.</mixed-citation></ref>
      <ref id="bib1.bib50"><label>50</label><?label 1?><mixed-citation>Klein Goldewijk, K., Dekker, S. C., and van Zanden, J. L.: Per-capita
estimations of long-term historical land use and the consequences for global
change research, J. Land Use Sci., 12, 313–337, <ext-link xlink:href="https://doi.org/10.1080/1747423X.2017.1354938" ext-link-type="DOI">10.1080/1747423X.2017.1354938</ext-link>, 2017b.</mixed-citation></ref>
      <ref id="bib1.bib51"><label>51</label><?label 1?><mixed-citation>Lawrence, D. M., Hurtt, G. C., Arneth, A., Brovkin, V., Calvin, K. V., Jones,
A. D., Jones, C. D., Lawrence, P. J., de Noblet-Ducoudré, N., Pongratz, J., Seneviratne, S. I., and Shevliakova, E.: The Land Use Model Intercomparison Project (LUMIP) contribution to CMIP6: rationale and experimental design, Geosci. Model Dev., 9, 2973–2998, <ext-link xlink:href="https://doi.org/10.5194/gmd-9-2973-2016" ext-link-type="DOI">10.5194/gmd-9-2973-2016</ext-link>, 2016.</mixed-citation></ref>
      <ref id="bib1.bib52"><label>52</label><?label 1?><mixed-citation>Le Quéré, C., Andrew, R. M., Friedlingstein, P., Sitch, S., Hauck, J., Pongratz, J., Pickers, P. A., Korsbakken, J. I., Peters, G. P., Canadell,
J. G., Arneth, A., Arora, V. K., Barbero, L., Bastos, A., Bopp, L., Chevallier, F., Chini, L. P., Ciais, P., Doney, S. C., Gkritzalis, T., Goll,
D. S., Harris, I., Haverd, V., Hoffman, F. M., Hoppema, M., Houghton, R. A.,
Hurtt, G., Ilyina, T., Jain, A. K., Johannessen, T., Jones, C. D., Kato, E.,
Keeling, R. F., Goldewijk, K. K., Landschützer, P., Lefèvre, N., Lienert, S., Liu, Z., Lombardozzi, D., Metzl, N., Munro, D .R., Nabel, J. E. M. S., Nakaoka, S.-I., Neill, C., Olsen, A., Ono, T., Patra, P., Peregon, A., Peters, W., Peylin, P., Pfeil, B., Pierrot, D., Poulter, B., Rehder, G.,
Resplandy, L., Robertson, E., Rocher, M., Rödenbeck, C., Schuster, U.,
Schwinger, J., Séférian, R., Skjelvan, I., Steinhoff, T., Sutton, A., Tans, P. P., Tian, H., Tilbrook, B., Tubiello, F. N., van der Laan-Luijkx, I. T., van der Werf, G. R., Viovy, N., Walker, A. P., Wiltshire, A. J., Wright, R., Zaehle, S., and Zheng, B.: Global Carbon Budget 2018, Earth Syst. Sci. Data, 10, 2141–2194, <ext-link xlink:href="https://doi.org/10.5194/essd-10-2141-2018" ext-link-type="DOI">10.5194/essd-10-2141-2018</ext-link>, 2018.</mixed-citation></ref>
      <ref id="bib1.bib53"><label>53</label><?label 1?><mixed-citation>Lewin, S. and Whitehouse, N.: Regional Land Use mapping code and dataset for Early and Middle Neolithic Ireland (3750–3600 cal BC and 3600–3400 cal BC, respectively), <ext-link xlink:href="https://doi.org/10.5281/zenodo.3625226" ext-link-type="DOI">10.5281/zenodo.3625226</ext-link>, 2020.</mixed-citation></ref>
      <ref id="bib1.bib54"><label>54</label><?label 1?><mixed-citation>
Loisel, J., Yu, Z., Beilman, D. W., Camill, P., Alm, J., Amesbury, M. J.,
Anderson, D., Andersson, S., Bochicchio, C., Barber, K., Belyea, L. R., Bunbury, J., Chambers, F. M., Charman, D. J., Vleeschouwer, F.D., Fiałkiewicz-Kozieł, B., Finkelstein, S. A., Gałka, M., Garneau, M.,
Hammarlund, D., Hinchcliffe, W., Holmquist, J., Hughes, P., Jones, M. C.,
Klein, E. S., Kokfelt, U., Korhola, A., Kuhry, P., Lamarre, A., Lamentowicz,
M., Large, D., Lavoie, M., MacDonald, G., Magnan, G., Mäkilä, M.,
Mallon, G., Mathijssen, P., Mauquoy, D., McCarroll, J., Moore, T. R., Nichols, J., O'Reilly, B., Oksanen, P., Packalen, M., Peteet, D., Richard,
P. J., Robinson, S., Ronkainen, T., Rundgren, M., Sannel, A. B. K., Tarnocai,
C., Thom, T., Tuittila, E.-S., Turetsky, M., Väliranta, M., and der Linden, M., van Geel B., van Bellen, S., Vitt, D., Zhao, Y., and Zhou, W.: A
database and synthesis of northern peatland soil properties and Holocene carbon and nitrogen accumulation, Holocene, 24, 1028–1042, 2014.</mixed-citation></ref>
      <ref id="bib1.bib55"><label>55</label><?label 1?><mixed-citation>
Lyman, R. L.: Quantitative Paleozoology, Cambridge University Press, Cambridge, 2008.</mixed-citation></ref>
      <ref id="bib1.bib56"><label>56</label><?label 1?><mixed-citation>Maezumi, S. Y., Robinson, M., de Souza, J., Urrego, D. H., Schaan, D., Alves,
D., and Iriarte, J.: New insights from pre-Columbian land use and fire management in Amazonian Dark Earth forests, Front. Ecol. Evol., 6, 111, <ext-link xlink:href="https://doi.org/10.3389/fevo.2018.00111" ext-link-type="DOI">10.3389/fevo.2018.00111</ext-link>, 2018.</mixed-citation></ref>
      <ref id="bib1.bib57"><label>57</label><?label 1?><mixed-citation>Mahowald, N. M., Randerson, J. T., Lindsay, K., Munoz, E., Doney, S. C.,
Lawrence, P., Schlunegger, S., Ward, D. S., Lawrence, D., and Hoffman, F. M.: Interactions between land use change and carbon cycle feedbacks, Global
Biogeochem. Cy., 31, 96–113, <ext-link xlink:href="https://doi.org/10.1002/2016GB005374" ext-link-type="DOI">10.1002/2016GB005374</ext-link>, 2017.</mixed-citation></ref>
      <ref id="bib1.bib58"><label>58</label><?label 1?><mixed-citation>
Marston, J. M.: Modeling wood aquisition strategies from archaeological charcoal remains, J. Arch. Sci., 36, 2192–2200, 2009.</mixed-citation></ref>
      <?pagebreak page822?><ref id="bib1.bib59"><label>59</label><?label 1?><mixed-citation>
Marston, J. M., Holdaway, S. J., and Wendrich, W.: Early- and middle-Holocene
wood exploitation in the Fayum basin, Egypt, Holocene, 27, 1812–1824, 2017.</mixed-citation></ref>
      <ref id="bib1.bib60"><label>60</label><?label 1?><mixed-citation>Mauri, A., Davis, B. A. S., Collins, P. M., and Kaplan, J. O.: The influence
of atmospheric circulation on the mid-Holocene climate of Europe: a data–model comparison, Clim. Past, 10, 1925–1938,
<ext-link xlink:href="https://doi.org/10.5194/cp-10-1925-2014" ext-link-type="DOI">10.5194/cp-10-1925-2014</ext-link>, 2014.</mixed-citation></ref>
      <ref id="bib1.bib61"><label>61</label><?label 1?><mixed-citation>
Mazoyer, M. and Roudart, L.: A History of World Agriculture: From the Neolithic to the Current Crisis, Earthscan, UK, 2006.</mixed-citation></ref>
      <ref id="bib1.bib62"><label>62</label><?label 1?><mixed-citation>McGrath, M. J., Luyssaert, S., Meyfroidt, P., Kaplan, J. O., Burgi, M., Chen, Y., Erb, K., Gimmi, U., McInerney, D., Naudts, K., Otto, J., Pasztor, F., Ryder, J., Schelhaas, M. J., and Valade, A.: Reconstructing European forest management from 1600 to 2010, Biogeoscience, 12, 4291–4316,
<ext-link xlink:href="https://doi.org/10.5194/bg-12-4291-2015" ext-link-type="DOI">10.5194/bg-12-4291-2015</ext-link>, 2015.</mixed-citation></ref>
      <ref id="bib1.bib63"><label>63</label><?label 1?><mixed-citation>McLaughlin, T. R., Whitehouse, N. J., Schulting, R. J., McClatchie, M., Barratt, P., and Bogaard, A.: The changing face of Neolithic and Bronze Age
Ireland: A big data approach to the settlement and burial records, J. World
Prehist., 29, 117–153, <ext-link xlink:href="https://doi.org/10.1007/s10963-016-9093-0" ext-link-type="DOI">10.1007/s10963-016-9093-0</ext-link>, 2016.</mixed-citation></ref>
      <ref id="bib1.bib64"><label>64</label><?label 1?><mixed-citation>Messori, G., Gaetani, M., Zhang, Q., Zhang, Q., and Pausata, F. S. R.: The
water cycle of the mid-Holocene West African monsoon: The role of vegetation
and dust emission changes, Int. J. Climatol., 39, 1927–1939, <ext-link xlink:href="https://doi.org/10.1002/joc.5924" ext-link-type="DOI">10.1002/joc.5924</ext-link>, 2019.</mixed-citation></ref>
      <ref id="bib1.bib65"><label>65</label><?label 1?><mixed-citation>Mitchell, L., Brook, E., Lee, J., Buizert, C., and Sowers, T.: Constraints on the late Holocene anthropogenic contribution to the atmospheric methane budget, Science, 342, 964–966, <ext-link xlink:href="https://doi.org/10.1126/science.1238920" ext-link-type="DOI">10.1126/science.1238920</ext-link>, 2013.</mixed-citation></ref>
      <ref id="bib1.bib66"><label>66</label><?label 1?><mixed-citation>
Morrison, K. D., Hammer, E., Popova, L., Madella, M., Whitehouse, N., Gaillard, M.-J., and LandCover6k Land-Use Group Members: Global-scale
comparisons of human land use: developing shared terminology for land-use
practices for global changes, PAGES Mag., 26, 8–9, 2018.</mixed-citation></ref>
      <ref id="bib1.bib67"><label>67</label><?label 1?><mixed-citation>
Myhre, G., Shindell, D., Bréon, F.-M., Collins, W., Fuglestvedt, J., Huang, J., Koch, D., Lamarque, J.-F., Lee, D., Mendoza, B., Nakajima, T.,
Robock, A., Stephens, G., Takemura T., and Zhang, H.: Anthropogenic and
natural radiative forcing, in: Climate Change 2013: The Physical Science
Basis. Contribution of Working Group I to the Fifth Assessment Report of the
Intergovernmental Panel on Climate Change, edited by: Stocker, T. F., Qin, D., Plattner, G.-K., Tignor, M., Allen, S. K., Boschung, J., Nauels, A., Xia, Y., Bex, V., and Midgley, P. M., Cambridge University Press Cambridge, Cambridge, UK and New York, NY, USA, 2013.</mixed-citation></ref>
      <ref id="bib1.bib68"><label>68</label><?label 1?><mixed-citation>
Nakagawa, T., Tarasov, P. E., Nishida, K., Gotanda, K., and Yasuda, Y.:
Quantitative pollen-based climate reconstruction in central Japan: application to surface and Late Quaternary spectra, Quaternary Sci. Rev., 21,
2099–2113, 2002.</mixed-citation></ref>
      <ref id="bib1.bib69"><label>69</label><?label 1?><mixed-citation>
Nielsen, A. B., Giesecke, T., Theuerkauf, M., Feeser, I., Behre, K.-H., Beug,
H.-J., Chen, S.-H., Christiansen, J., Dörfler, W., Endtmann, E., Jahns, S., de Klerk, O., Kühl, N., Latałowa, M., Odgaard, B. V., Rasmussen,
P., Stockholm, J. R., Voigt, R., Wiethold, J., and Wolters, S.: Quantitative
reconstructions of changes in regional openness in north-central Europe
reveal new insights into old questions, Quaternary Sci. Rev., 47, 131–149, 2012.</mixed-citation></ref>
      <ref id="bib1.bib70"><label>70</label><?label 1?><mixed-citation>Oh, Y., Conte, M., Kang, S., Kim, J., and Hwang, J.: Population fluctuation
and the adoption of food production in prehistoric Korea: using radiocarbon
dates as a proxy for population change, Radiocarbon, 59, 1761–1770, <ext-link xlink:href="https://doi.org/10.1017/RDC.2017.122" ext-link-type="DOI">10.1017/RDC.2017.122</ext-link>, 2017.</mixed-citation></ref>
      <ref id="bib1.bib71"><label>71</label><?label 1?><mixed-citation>
Ortega-Rosas, C. I., Guiot, J., Penalba, M. C., and Ortiz-Acosta, M. E.:
Biomization and quantitative climate reconstruction techniques in northwestern Mexico – with an application to four Holocene pollen sequences,
Global Planet. Change, 61, 242–266, 2008.</mixed-citation></ref>
      <ref id="bib1.bib72"><label>72</label><?label 1?><mixed-citation>Orton, D., Gaastra, J., and Vander Linden, M.: Between the Danube and the Deep Blue Sea: zooarchaeological meta-analysis reveals variability in the
spread and development of Neolithic farming across the western Balkans, Open
Quat., 2, 6, <ext-link xlink:href="https://doi.org/10.5334/oq.28" ext-link-type="DOI">10.5334/oq.28</ext-link>, 2016.</mixed-citation></ref>
      <ref id="bib1.bib73"><label>73</label><?label 1?><mixed-citation>Otto-Bliesner, B. L., Braconnot, P., Harrison, S. P., Lunt, D. J., Abe-Ouchi, A., Albani, S., Bartlein, P. J., Capron, E., Carlson, A. E., Dutton, A., Fischer, H., Goelzer, H., Govin, A., Haywood, A., Joos, F., LeGrande, A. N., Lipscomb, W. H., Lohmann, G., Mahowald, N., Nehrbass-Ahles, C., Pausata, F. S. R., Peterschmitt, J.-Y., Phipps, S. J., Renssen, H., and Zhang, Q.: The PMIP4 contribution to CMIP6 – Part 2: Two interglacials, scientific objective and experimental design for Holocene and Last Interglacial simulations, Geosci. Model Dev., 10, 3979–4003, <ext-link xlink:href="https://doi.org/10.5194/gmd-10-3979-2017" ext-link-type="DOI">10.5194/gmd-10-3979-2017</ext-link>, 2017.</mixed-citation></ref>
      <ref id="bib1.bib74"><label>74</label><?label 1?><mixed-citation>Pausata, F. S. R., Messori, G., and Zhang, Q.: Impacts of dust reduction on the northward expansion of the African monsoon during the Green Sahara period, Earth Planet. Sc. Lett., 434, 298–307, <ext-link xlink:href="https://doi.org/10.1016/j.epsl.2015.11.049" ext-link-type="DOI">10.1016/j.epsl.2015.11.049</ext-link>, 2016.</mixed-citation></ref>
      <ref id="bib1.bib75"><label>75</label><?label 1?><mixed-citation>Perugini, L. Caporaso, L., Marconi, S., Cescatti, A., Quesada, B., de Noblet-Ducoudré, N., House, J. I., and Arneth, A.: Biophysical effects on temperature and precipitation due to land cover change, Environ. Res. Lett., 12, 053002, <ext-link xlink:href="https://doi.org/10.1088/1748-9326/aa6b3f" ext-link-type="DOI">10.1088/1748-9326/aa6b3f</ext-link>, 2017.</mixed-citation></ref>
      <ref id="bib1.bib76"><label>76</label><?label 1?><mixed-citation>Phelps, L. N. and Kaplan, J. O.: Land use for animal production in global
change studies: Defining and characterizing a framework, Global Change Biol.,
23, 4457–4471, <ext-link xlink:href="https://doi.org/10.1111/gcb.13732" ext-link-type="DOI">10.1111/gcb.13732</ext-link>, 2017.</mixed-citation></ref>
      <ref id="bib1.bib77"><label>77</label><?label 1?><mixed-citation>
Pirzamanbein, B., Lindström, J., Poska, A., Sugita, S., Trondman, A.,
Fyfe, R., Mazier, F., Nielsen, A. B., Kaplan, J. O., Bjune, A. E., Birks, H. J. B., Giesecke, T., Kangur, M., Latałowa, M., Marquer, L., Smith, B., and Gaillard, M.-J.: Creating spatially continuous maps of past land cover from point estimates: A new statistical approach applied to pollen data, Ecol. Complex., 20, 127–141, 2014.</mixed-citation></ref>
      <ref id="bib1.bib78"><label>78</label><?label 1?><mixed-citation>
Pirzamanbein, B., Lindström, J., Poska, A., and Gaillard, M.-J.: Modelling spatial compositional data: Reconstructions of past land cover and
uncertainties, Spat. Stat., 24, 14–31, 2018.</mixed-citation></ref>
      <ref id="bib1.bib79"><label>79</label><?label 1?><mixed-citation>Pongratz, J., Reick, C., Raddatz, T., and Claussen, M.: A reconstruction of
global agricultural areas and land cover for the last millennium, Global
Biogeochem. Cy., 22, GB3018, <ext-link xlink:href="https://doi.org/10.1029/2007GB003153" ext-link-type="DOI">10.1029/2007GB003153</ext-link>, 2008.</mixed-citation></ref>
      <ref id="bib1.bib80"><label>80</label><?label 1?><mixed-citation>Pongratz, J., Reick, C. H., Raddatz, T., and Claussen, M.: Biogeophysical
versus biogeochemical climate response to historical anthropogenic land cover change, Geophys. Res. Lett., 37, L08702, <ext-link xlink:href="https://doi.org/10.1029/2010GL043010" ext-link-type="DOI">10.1029/2010GL043010</ext-link>, 2010.</mixed-citation></ref>
      <ref id="bib1.bib81"><label>81</label><?label 1?><mixed-citation>Pongratz, J., Reick, C. H., Houghton, R. A., and House, J. I.: Terminology as a key uncertainty in net land use and land cover change carbon flux estimates, Earth Syst. Dynam., 5, 177–195, <ext-link xlink:href="https://doi.org/10.5194/esd-5-177-2014" ext-link-type="DOI">10.5194/esd-5-177-2014</ext-link>, 2014.</mixed-citation></ref>
      <?pagebreak page823?><ref id="bib1.bib82"><label>82</label><?label 1?><mixed-citation>
Ramankutty, N. and Foley, J. A.: Estimating historical changes in global land cover: Croplands from 1700 to 1992, Global Biogeochem. Cy., 13, 997–1027, 1999.</mixed-citation></ref>
      <ref id="bib1.bib83"><label>83</label><?label 1?><mixed-citation>
Reitz, E. J. and Wing, E. S.: Zooarchaeology, Cambridge University Press,
Cambridge, 2008.</mixed-citation></ref>
      <ref id="bib1.bib84"><label>84</label><?label 1?><mixed-citation>
Rick, J. W.: Dates as data: an examination of the Peruvian Preceramic
radiocarbon record, Am. Antiq., 52, 55–73, 1987.</mixed-citation></ref>
      <ref id="bib1.bib85"><label>85</label><?label 1?><mixed-citation>
Robinson, E., Zahid, H. J., Codding, B. F., Haas, R., and Kelly, R. L.:
Spatiotemporal dynamics of prehistoric human population growth: radiocarbon
`dates as data' and population ecology models, J. Arch. Sci., 101, 63–71,
2019.</mixed-citation></ref>
      <ref id="bib1.bib86"><label>86</label><?label 1?><mixed-citation>Ruddiman, W. F.: The anthropogenic greenhouse era began thousands of years ago, Climatic Change, 61, 261–293, <ext-link xlink:href="https://doi.org/10.1023/B:CLIM.0000004577.17928.fa" ext-link-type="DOI">10.1023/B:CLIM.0000004577.17928.fa</ext-link>,
2003.</mixed-citation></ref>
      <ref id="bib1.bib87"><label>87</label><?label 1?><mixed-citation>Russell, T., Silva, F., and Steele, J.: Modelling the spread of farming in
the Bantu-speaking regions of Africa: an archaeology-based phylogeography,
Plos One, 9, e87584, <ext-link xlink:href="https://doi.org/10.1371/journal.pone.0087854" ext-link-type="DOI">10.1371/journal.pone.0087854</ext-link>, 2014.</mixed-citation></ref>
      <ref id="bib1.bib88"><label>88</label><?label 1?><mixed-citation>Shennan, S., Downey, S. S., Timpson, A., Edinborough, K., Colledge, S., Kerig, T., Manning, K., and Thomas, M. G.: Regional population collapse followed initial agriculture booms in mid-Holocene Europe, Nat. Commun., 4,
248, <ext-link xlink:href="https://doi.org/10.1038/ncomms3486" ext-link-type="DOI">10.1038/ncomms3486</ext-link>, 2013.</mixed-citation></ref>
      <ref id="bib1.bib89"><label>89</label><?label 1?><mixed-citation>Shevliakova, E., Pacala, S. W., Malyshev, S., Hurtt, G. C., Milly, P. C. D.,
Caspersen, J. P., Sentman, L. T., Fisk, J. P., Wirth, C., and Crevoisier, C.: Carbon cycling under 300 years of land use change: Importance of the secondary vegetation sink, Global Biogeochem. Cy., 23, GB2022,
<ext-link xlink:href="https://doi.org/10.1029/2007GB003176" ext-link-type="DOI">10.1029/2007GB003176</ext-link>, 2009.</mixed-citation></ref>
      <ref id="bib1.bib90"><label>90</label><?label 1?><mixed-citation>Sigl, M., Winstrup, M., McConnell, J. R., Welten, K. C., Plunkett, G., Ludlow, F., Büntgen, U., Caffee, M., Chellman, N., Dahl-Jensen, D., Fischer, H., Kipfstuhl, S., Kostick, C., Maselli, O. J., Mekhaldi, F.,
Mulvaney, R., Muscheler, R., Pasteris, D. R., Pilcher, J. R., Salzer, M.,
Schüpbach, S., Steffensen, J. P., Vinther, B. M., and Woodruff, T. E.: Timing and climate forcing of volcanic eruptions for the past 2,500 years, Nature, 523, 543–549, <ext-link xlink:href="https://doi.org/10.1038/nature14565" ext-link-type="DOI">10.1038/nature14565</ext-link>, 2015.</mixed-citation></ref>
      <ref id="bib1.bib91"><label>91</label><?label 1?><mixed-citation>Silva, F. and Steele, J.: New methods for reconstructing geographical effects on dispersal rates from large-scale radiocarbon databases, J. Arch. Sci., 52, 609–620, <ext-link xlink:href="https://doi.org/10.1016/j.jas.2014.04.021" ext-link-type="DOI">10.1016/j.jas.2014.04.021</ext-link>, 2014.</mixed-citation></ref>
      <ref id="bib1.bib92"><label>92</label><?label 1?><mixed-citation>Silva, F. and Vander Linden, M.: Amplitude of travelling front as inferred
from <inline-formula><mml:math id="M45" display="inline"><mml:mrow class="chem"><mml:msup><mml:mi/><mml:mn mathvariant="normal">14</mml:mn></mml:msup><mml:mi mathvariant="normal">C</mml:mi></mml:mrow></mml:math></inline-formula> predicts levels of genetic admixture among European early
farmers, Sci. Rep., 7, 11985, <ext-link xlink:href="https://doi.org/10.1038/s41598-017-12318-2" ext-link-type="DOI">10.1038/s41598-017-12318-2</ext-link>, 2017.</mixed-citation></ref>
      <ref id="bib1.bib93"><label>93</label><?label 1?><mixed-citation>Silva, F., Stevens, C. J., Weisskopf, A., Castillo, C., Qin, L., Bevan, A.,
and Fuller, D. Q.: Modelling the geographical origin of rice cultivation in
Asia using the Rice Archaeological Database, Plos One, 10, e0137024, <ext-link xlink:href="https://doi.org/10.1371/journal.pone.0137024" ext-link-type="DOI">10.1371/journal.pone.0137024</ext-link>, 2015.</mixed-citation></ref>
      <ref id="bib1.bib94"><label>94</label><?label 1?><mixed-citation>Singarayer, J. S., Valdes, P. J., Friedlingstein, P., Nelson, S., and Beerling, D. J.: Late Holocene methane rise caused by orbitally controlled
increase in tropical sources, Nature, 470, 82–85, <ext-link xlink:href="https://doi.org/10.1038/nature09739" ext-link-type="DOI">10.1038/nature09739</ext-link>,
2011.</mixed-citation></ref>
      <ref id="bib1.bib95"><label>95</label><?label 1?><mixed-citation>Smith, M. C., Singarayer, J. S., Valdes, P. J., Kaplan, J. O., and Branch, N. P.: The biogeophysical climatic impacts of anthropogenic land use change during the Holocene, Clim. Past, 12, 923–941, <ext-link xlink:href="https://doi.org/10.5194/cp-12-923-2016" ext-link-type="DOI">10.5194/cp-12-923-2016</ext-link>, 2016.</mixed-citation></ref>
      <ref id="bib1.bib96"><label>96</label><?label 1?><mixed-citation>
Steinhilber, F., Abreu, J. A., Beer, J., Brunner, I., Christl, M., Fischer,
H., Heikkilä, U., Kubik, P. W., Mann, M., McCracken, K. G., Miller, H.,
Miyahara, H., Oerter, H., and Wilhelms, F.: 9400 years of cosmic radiation
and solar activity from ice cores and tree rings, P. Natl. Acad. Sci. USA, 109, 5967–5971, 2012.</mixed-citation></ref>
      <ref id="bib1.bib97"><label>97</label><?label 1?><mixed-citation>Stocker, B. D., Strassmann, K., and Joos, F.: Sensitivity of Holocene atmospheric <inline-formula><mml:math id="M46" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> and the modern carbon budget to early human land use: analyses with a process-based model, Biogeosciences, 8, 69–88,
<ext-link xlink:href="https://doi.org/10.5194/bg-8-69-2011" ext-link-type="DOI">10.5194/bg-8-69-2011</ext-link>, 2011.</mixed-citation></ref>
      <ref id="bib1.bib98"><label>98</label><?label 1?><mixed-citation>Stocker, B. D., Yu, Z., Massa, C., and Joos, F.: Holocene peatland and
ice-core data constraints on the timing and magnitude of <inline-formula><mml:math id="M47" display="inline"><mml:mrow class="chem"><mml:msub><mml:mi mathvariant="normal">CO</mml:mi><mml:mn mathvariant="normal">2</mml:mn></mml:msub></mml:mrow></mml:math></inline-formula> emissions from past land use, P. Natl. Acad. Sci. USA, 114, 1492–1497,
<ext-link xlink:href="https://doi.org/10.1073/pnas.1613889114" ext-link-type="DOI">10.1073/pnas.1613889114</ext-link>, 2017.</mixed-citation></ref>
      <ref id="bib1.bib99"><label>99</label><?label 1?><mixed-citation>Styring, A., Rösch, M., Stephan, E., Stika, H.-P., Fischer, E., Sillmann, E., and Bogaard, A.: Centralisation and long-term change in farming regimes: comparing agricultural practices in Neolithic and Iron Age south-west Germany, Proc. Prehist. Soc., 83, 357–381, <ext-link xlink:href="https://doi.org/10.1017/ppr.2017.3" ext-link-type="DOI">10.1017/ppr.2017.3</ext-link>, 2017.</mixed-citation></ref>
      <ref id="bib1.bib100"><label>100</label><?label 1?><mixed-citation>
Sugita, S.: Theory of quantitative reconstruction of vegetation I: pollen from large sites REVEALS regional vegetation composition, Holocene, 17,
229–241, 2007.</mixed-citation></ref>
      <ref id="bib1.bib101"><label>101</label><?label 1?><mixed-citation>
Tarasov, P., Williams, J. W., Andreev, A., Nakagawa, T., Bezrukova, E.,
Herzschuh, U., Igarashi, Y., Müller, S., Werner, K., and Zheng, Z.:
Satellite- and pollen-based quantitative woody cover reconstructions for
northern Asia: Verification and application to late-Quaternary pollen data,
Earth Planet. Sc. Lett., 264, 284–298, 2007.</mixed-citation></ref>
      <ref id="bib1.bib102"><label>102</label><?label 1?><mixed-citation>
Tauger, M. B.: Agriculture in World History, Routledge, Oxford, 2013.</mixed-citation></ref>
      <ref id="bib1.bib103"><label>103</label><?label 1?><mixed-citation>Tierney, J. E., Pausata, F. S. R., and deMenocal, P. B.: Rainfall regimes of the Green Sahara, Sci. Adv., 3, e1601503, <ext-link xlink:href="https://doi.org/10.1126/sciadv.1601503" ext-link-type="DOI">10.1126/sciadv.1601503</ext-link>, 2017.</mixed-citation></ref>
      <ref id="bib1.bib104"><label>104</label><?label 1?><mixed-citation>Timpson, A., Colledge, S., Crema, E., Edinborough, K., Kerig, T., Manning, K., Thomas, M. G., and Shennan, S.: Reconstructing regional population
fluctuations in the European Neolithic using radiocarbon dates: a new case-study using an improved method, J. Arch. Sci., 52, 549–557,
<ext-link xlink:href="https://doi.org/10.1016/j.jas.2014.08.011" ext-link-type="DOI">10.1016/j.jas.2014.08.011</ext-link>, 2014.</mixed-citation></ref>
      <ref id="bib1.bib105"><label>105</label><?label 1?><mixed-citation>Toohey, M. and Sigl, M.: Volcanic stratospheric sulphur injections and
aerosol optical depth from 500 BCE to 1900 CE, Earth Syst. Sci. Data, 9,
809–831, <ext-link xlink:href="https://doi.org/10.5194/essd-9-809-2017" ext-link-type="DOI">10.5194/essd-9-809-2017</ext-link>, 2017.</mixed-citation></ref>
      <ref id="bib1.bib106"><label>106</label><?label 1?><mixed-citation>Trondman, A. K., Gaillard, M.-J., Mazier, F., Sugita, S., Fyfe, R., Nielsen,
A. B., Twiddle, C., Barratt, P., Birks, H. J. B., Bjune, A. E., Björkman,
L., Broström, A., Caseldine, C., David, R., Dodson, J., Dörfler, W.,
Fischer, E., van Geel, B., Giesecke, T., Hultberg, T., Kalnina, L., Kangur,
M., van der Knaap, P., Koff, T., Kuneš, P., Lagerås, P., Latałowa,
M., Lechterbeck, J., Leroyer, C., Leydet, M., Lindbladh, M., Marquer, L.,
Mitchell, F. J. G., Odgaard, B. V., Peglar, S. M., Persson, T., Poska, A.,
Rösch, M., Seppä, H., Veski, S., and Wick, L.: Pollen-based quantitative reconstructions of Holocene regional vegetation cover (plant-functional types and land-cover types) in Europe suitable for climate
modelling, Global Change Biol., 21, 676–697, <ext-link xlink:href="https://doi.org/10.1111/gcb.12737" ext-link-type="DOI">10.1111/gcb.12737</ext-link>, 2015.</mixed-citation></ref>
      <ref id="bib1.bib107"><label>107</label><?label 1?><mixed-citation>Trondman, A.-K., Gaillard, M.-J., Sugita, S., Björkman, L., Greisman, A., Hultberg, T., Lagerås, P., and Lindbladh, M.: Are pollen records from small sites appropriate for REVEALS model-based quantitative reconstructions of past regional vegetation? A<?pagebreak page824?>n empirical test in southern Sweden, Veget. Hist. Archaeobot., 25, 131–151, <ext-link xlink:href="https://doi.org/10.1007/s00334-015-0536-9" ext-link-type="DOI">10.1007/s00334-015-0536-9</ext-link>, 2016.</mixed-citation></ref>
      <ref id="bib1.bib108"><label>108</label><?label 1?><mixed-citation>van den Hurk, B., Kim, H., Krinner, G., Seneviratne, S. I., Derksen, C., Oki,
T., Douville, H., Colin, J., Ducharne, A., Cheruy, F., Viovy, N., Puma, M. J., Wada, Y., Li, W., Jia, B., Alessandri, A., Lawrence, D. M., Weedon, G. P., Ellis, R., Hagemann, S., Mao, J., Flanner, M. G., Zampieri, M., Materia, S., Law, R. M., and Sheffield, J.: LS3MIP (v1.0) contribution to CMIP6: the Land Surface, Snow and Soil moisture Model Intercomparison Project – aims, setup and expected outcome, Geosci. Model Dev., 9, 2809–2832, <ext-link xlink:href="https://doi.org/10.5194/gmd-9-2809-2016" ext-link-type="DOI">10.5194/gmd-9-2809-2016</ext-link>, 2016.</mixed-citation></ref>
      <ref id="bib1.bib109"><label>109</label><?label 1?><mixed-citation>Vander Linden, M.: Code for fig 3 of <ext-link xlink:href="https://doi.org/10.5194/gmd-2019-125" ext-link-type="DOI">10.5194/gmd-2019-125</ext-link>, available at: <uri>https://github.com/mavdlind/GMD</uri>, last access: 14 February 2020.</mixed-citation></ref>
      <ref id="bib1.bib110"><label>110</label><?label 1?><mixed-citation>
Vavrus, S., Ruddiman, W. F., and Kutzbach, J. E.: Climate model tests of the
anthropogenic influence on greenhouse-induced climate change: the role of
early human agriculture, industrialization, and vegetation feedbacks, Quaternary Sci. Rev., 27, 1410–1425, 2008.</mixed-citation></ref>
      <ref id="bib1.bib111"><label>111</label><?label 1?><mixed-citation>
Veal, R.: Wood and charcoal for Rome: towards an understanding of ancient regional fuel economics, in: Rural communities in a globalizing economy: new perspectives on the economic integration of Roman Italy, edited by: de Haas, T. and Gijs, T., Brill, New York, Leiden, 388–406, 2017.</mixed-citation></ref>
      <ref id="bib1.bib112"><label>112</label><?label 1?><mixed-citation>
Viau, A. E. and Gajewski, K.: Reconstructing millennial, regional paleoclimates of boreal Canada during the Holocene, J. Climate, 22, 316–330,
2009.</mixed-citation></ref>
      <ref id="bib1.bib113"><label>113</label><?label 1?><mixed-citation>Viau, A. E., Gajewski, K., Sawada, M., and Fines, P.: Mean-continental July temperature variability in North America during the past 14,000 years, J. Geophys. Res.-Atmos., 111, D09102, <ext-link xlink:href="https://doi.org/10.1029/2005JD006031" ext-link-type="DOI">10.1029/2005JD006031</ext-link>, 2006.</mixed-citation></ref>
      <ref id="bib1.bib114"><label>114</label><?label 1?><mixed-citation>Weiberg, E., Hughes, R. E., Finné, M., Bonnier, A., and Kaplan, J. O.:
Mediterranean land use systems from prehistory to antiquity: a case study
from Peloponnese (Greece), J. Land Use Sci., 14, 1–20, <ext-link xlink:href="https://doi.org/10.1080/1747423x.2019.1639836" ext-link-type="DOI">10.1080/1747423x.2019.1639836</ext-link>, 2019.
</mixed-citation></ref><?xmltex \hack{\newpage}?>
      <ref id="bib1.bib115"><label>115</label><?label 1?><mixed-citation>Whitehouse, N., Schulting, R. J., McClatchie, M., Barratt, P., LcLaughlin, T. R., Bogaard, A., Colledge, S., Marchant, R., Gaffrey, J., and Bunting, M. J.: Neolithic agriculture on the European western frontier: the boom and bust of early farming in Ireland, J. Arch. Sci., 51, 181–205, <ext-link xlink:href="https://doi.org/10.1016/j.jas.2013.08.009" ext-link-type="DOI">10.1016/j.jas.2013.08.009</ext-link>, 2014.</mixed-citation></ref>
      <ref id="bib1.bib116"><label>116</label><?label 1?><mixed-citation>Williams, A.: The use of summed radiocarbon probability distributions in
archaeology: a review of methods, J. Archaeol. Sci., 39, 578–589,
<ext-link xlink:href="https://doi.org/10.1016/j.jas.2011.07.014" ext-link-type="DOI">10.1016/j.jas.2011.07.014</ext-link>, 2012.</mixed-citation></ref>
      <ref id="bib1.bib117"><label>117</label><?label 1?><mixed-citation>
Wilmshurst, J. M., McGlone, M. S., Leathwick, J. R., and Newnham, R. M.: A
pre-deforestation pollen-climate calibration model for New Zealand and
quantitative temperature reconstructions for the past 18 000 years BP, J.
Quaternay Sci., 22, 535–547, 2007.</mixed-citation></ref>
      <ref id="bib1.bib118"><label>118</label><?label 1?><mixed-citation>Wright, P.: Preservation or destruction of plant remains by carbonization, J. Arch. Sci., 30, 577–583, <ext-link xlink:href="https://doi.org/10.1016/S0305-4403(02)00203-0" ext-link-type="DOI">10.1016/S0305-4403(02)00203-0</ext-link>, 2003.</mixed-citation></ref>
      <ref id="bib1.bib119"><label>119</label><?label 1?><mixed-citation>Zahid, H. J., Robinson, E., and Kelly, R. L.: Agriculture, population growth,
and statistical analysis of the radiocarbon record, P. Natl. Acad. Sci. USA,
113, 931–935, <ext-link xlink:href="https://doi.org/10.1073/pnas.1517650112" ext-link-type="DOI">10.1073/pnas.1517650112</ext-link>, 2016.</mixed-citation></ref>
      <ref id="bib1.bib120"><label>120</label><?label 1?><mixed-citation>Zanon, M., Davis, B. A. S., Marquer, L., Brewer, S., and Kaplan, J. O.: European forest cover during the past 12,000 years: a palynological reconstruction based on modern analogs and remote sensing, Front. Plant
Sci., 9, 253, <ext-link xlink:href="https://doi.org/10.3389/fpls.2018.00253" ext-link-type="DOI">10.3389/fpls.2018.00253</ext-link>, 2018.</mixed-citation></ref>
      <ref id="bib1.bib121"><label>121</label><?label 1?><mixed-citation>Zimmermann, A., Wendt, K. P., and Hilpert, J.: Landscape archaeology in
central Europe, Proc. Prehist. Soc., 75, 1–53, <ext-link xlink:href="https://doi.org/10.1017/S0079497X00000281" ext-link-type="DOI">10.1017/S0079497X00000281</ext-link>, 2009.</mixed-citation></ref>
      <ref id="bib1.bib122"><label>122</label><?label 1?><mixed-citation>
Zohary, D., Hopf, M., and Weiss, E.: Domestication of Plants in the Old World: The Origin and Spread of Domesticated Plants in South-west Asia,
Europe, and the Mediterranean Basin, 4th Edn., Oxford University Press, Oxford, 2012.</mixed-citation></ref>

  </ref-list></back>
    <!--<article-title-html>Development and testing scenarios for implementing  land use and land cover changes during the Holocene  in Earth system model experiments</article-title-html>
<abstract-html><p>Anthropogenic changes in land use and land cover (LULC) during the
pre-industrial Holocene could have affected regional and global climate.
Existing scenarios of LULC changes during the Holocene are based on
relatively simple assumptions and highly uncertain estimates of population
changes through time. Archaeological and palaeoenvironmental reconstructions have the potential to refine these assumptions and estimates. The Past Global Changes (PAGES) LandCover6k initiative is working towards improved reconstructions of LULC globally. In this paper, we document the types of archaeological data that are being collated and how they will be used to improve LULC reconstructions. Given the large methodological uncertainties involved, both in reconstructing LULC from the archaeological data and in implementing these reconstructions into global scenarios of LULC, we propose a protocol to evaluate the revised scenarios using independent pollen-based reconstructions of land cover and climate. Further evaluation of the revised scenarios involves carbon cycle model simulations to determine whether the LULC reconstructions are consistent with constraints provided by ice core records of CO<sub>2</sub> evolution and modern-day LULC. Finally, the protocol outlines how the improved LULC reconstructions will be used in palaeoclimate simulations in the Palaeoclimate Modelling Intercomparison Project to quantify the magnitude of anthropogenic impacts on climate through time and ultimately to improve the realism of Holocene climate simulations.</p></abstract-html>
<ref-html id="bib1.bib1"><label>1</label><mixed-citation>
Arneth, A., Sitch, S., Pongratz, J., Stocker, B. D., Ciais, P., Poulter, B.,
Bayer, A. D., Bondeau, A., Calle, L., Chini, L. P., Gasser, T., Fader, M.,
Friedlingstein, P., Kato, E., Li, W., Lindeskog, M., Nabel, J. E. M. S., Pugh, T. A. M., Robertson, E., Viovy, N., Yue, C., and Zaehle, S.: Historical carbon dioxide emissions caused by land-use changes are possibly larger than
assumed, Nat. Geosci., 10, 79–84, <a href="https://doi.org/10.1038/ngeo2882" target="_blank">https://doi.org/10.1038/ngeo2882</a>, 2017.
</mixed-citation></ref-html>
<ref-html id="bib1.bib2"><label>2</label><mixed-citation>
Arneth, A., Denton, F., Agus, F., Elbehri, A., Erb, K., Elasha, B. O., Rahimi, M., Rounsevell, M., Spence, A., and Valentini, R.: IPCC Special
Report on Climate Change, Desertification, Land Degradation, Sustainable
Land Management, Food Security, and Greenhouse gas fluxes in Terrestrial
Ecosystems, 2019.
</mixed-citation></ref-html>
<ref-html id="bib1.bib3"><label>3</label><mixed-citation>
Balsera, V., Díaz-del-Río, P., Gilman, A., Uriarte, A., and Vicent, J. M.: Approaching the demography of late prehistoric Iberia through summed calibrated probability distributions (7000–2000&thinsp;cal&thinsp;BC), Quatern. Int.,
208–211, <a href="https://doi.org/10.1016/j.quaint.2015.06.022" target="_blank">https://doi.org/10.1016/j.quaint.2015.06.022</a>, 2015.
</mixed-citation></ref-html>
<ref-html id="bib1.bib4"><label>4</label><mixed-citation>
Bartlein, P. J., Harrison, S. P., Brewer, S., Connor, S., Davis, B. A. S.,
Gajewski, K., Guiot, J., Harrison-Prentice, T. I., Henderson, A., Peyron, O., Prentice, I. C., Scholze, M., Seppä, H., Shuman, B., Sugita, S., Thompson, R. S., Viau, A., Williams, J., and Wu, H.: Pollen-based continental climate reconstructions at 6 and 21&thinsp;ka: a global synthesis, Clim. Dynam., 37, 775–802, <a href="https://doi.org/10.1007/s00382-010-0904-1" target="_blank">https://doi.org/10.1007/s00382-010-0904-1</a>, 2011.
</mixed-citation></ref-html>
<ref-html id="bib1.bib5"><label>5</label><mixed-citation>
Bartlein, P. J., Harrison, S. P., and Izumi, K.: Underlying causes of Eurasian mid-continental aridity in simulations of mid-Holocene climate, Geophys. Res. Lett., 44, 9020–9028, <a href="https://doi.org/10.1002/2017GL074476" target="_blank">https://doi.org/10.1002/2017GL074476</a>, 2017.
</mixed-citation></ref-html>
<ref-html id="bib1.bib6"><label>6</label><mixed-citation>
Barton, C. M., Ullah, I. I., and Bergin, S.: Land use, water and Mediterranean landscapes: modelling long-term dynamics of complex socio-ecological systems, Philos. T. Roy. Soc. A,368, 5275–5297, <a href="https://doi.org/10.1098/rsta.2010.0193" target="_blank">https://doi.org/10.1098/rsta.2010.0193</a>, 2010.
</mixed-citation></ref-html>
<ref-html id="bib1.bib7"><label>7</label><mixed-citation>
Berger, A.  and Loutre, M.-F.: Insolation values for the climate of the last
10 million of years, Quaternary Sci. Rev., 10, 297–317, <a href="https://doi.org/10.1016/0277-3791(91)90033-Q" target="_blank">https://doi.org/10.1016/0277-3791(91)90033-Q</a>, 1991.
</mixed-citation></ref-html>
<ref-html id="bib1.bib8"><label>8</label><mixed-citation>
Bishop, R. R., Church, M. J., and Rowley-Conwy, P. A.: Firewood, food and human niche construction: the potential role of Mesolithic hunter–gatherers in actively structuring Scotland's woodlands, Quaternary Sci. Rev., 108, 51–75, 2015.
</mixed-citation></ref-html>
<ref-html id="bib1.bib9"><label>9</label><mixed-citation>
Bocquet-Appel, J.-P., Naji, S., Vander Linden, M., and Kozłowsi, J. K.:
Detection of diffusion and contact zones of early farming in Europe from the
space-time distribution of <sup>14</sup>C dates, J. Arch. Sci., 36, 807–820,
<a href="https://doi.org/10.1016/j.jas.2008.11.004" target="_blank">https://doi.org/10.1016/j.jas.2008.11.004</a>, 2009.
</mixed-citation></ref-html>
<ref-html id="bib1.bib10"><label>10</label><mixed-citation>
Crema, E. R., Habu, J., Kobayashi, K., and Madella, M.: Summed probability
distribution of <sup>14</sup>C dates suggests regional divergences in the
population dynamics of the Jomon period in eastern Japan, PlosOne, 11,
e0154809, <a href="https://doi.org/10.1371/journal.pone.0154809" target="_blank">https://doi.org/10.1371/journal.pone.0154809</a>, 2016.
</mixed-citation></ref-html>
<ref-html id="bib1.bib11"><label>11</label><mixed-citation>
Davis, B. A. S., Brewer, S., Stevenson, A. C., Guiot, J., and Juggins, S.: The temperature of Europe during the Holocene reconstructed from pollen data,
Quaternary Sci. Rev., 22, 1701–1716, 2003.
</mixed-citation></ref-html>
<ref-html id="bib1.bib12"><label>12</label><mixed-citation>
Davison, K., Dolukhanov, P., Sarson, G. R., and Shukurov, P.: The role of
waterways in the spread of the Neolithic, J. Arch. Sci., 33, 641–652, <a href="https://doi.org/10.1016/j.jas.2005.09.017" target="_blank">https://doi.org/10.1016/j.jas.2005.09.017</a>, 2006.
</mixed-citation></ref-html>
<ref-html id="bib1.bib13"><label>13</label><mixed-citation>
Dawson, A.: Code to reproduce Figure 7 from Harrison et al., GMD, <a href="https://doi.org/10.5281/zenodo.3604328" target="_blank">https://doi.org/10.5281/zenodo.3604328</a>, 2019.
</mixed-citation></ref-html>
<ref-html id="bib1.bib14"><label>14</label><mixed-citation>
Dawson, A., Paciorek, C. J., McLachlan, J. S., Goring, S., Williams, J. W., and Jackson, S. T.: Quantifying pollen-vegetation relationships to reconstruct ancient forests using 19th-century forest composition and pollen data, Quaternary Sci. Rev., 137, 156–175, <a href="https://doi.org/10.1016/j.quascirev.2016.01.012" target="_blank">https://doi.org/10.1016/j.quascirev.2016.01.012</a>, 2016.
</mixed-citation></ref-html>
<ref-html id="bib1.bib15"><label>15</label><mixed-citation>
Dawson, A., Cao, X., Chaput, M., Hopla, E., Li, F., Edwards, M., Fyfe, R.,
Gajewski, K., Goring, S. J., Herzschuh, U., Mazier, F., Sugita, S., Williams,
J. W., Xu, Q., and Gaillard, M.-J.: Finding the magnitude of human induced
Northern Hemisphere land-cover transformation between 6 and 0.2&thinsp;ka&thinsp;BP, PAGES Mag., 26, 34–35, <a href="https://doi.org/10.22498/pages.26.1.34" target="_blank">https://doi.org/10.22498/pages.26.1.34</a>, 2018.
</mixed-citation></ref-html>
<ref-html id="bib1.bib16"><label>16</label><mixed-citation>
Egerer, S., Claussen, M., and Reick, C.: Rapid increase in simulated North
Atlantic dust deposition due to fast change of northwest African landscape
during the Holocene, Clim. Past, 14, 1051–1066,
<a href="https://doi.org/10.5194/cp-14-1051-2018" target="_blank">https://doi.org/10.5194/cp-14-1051-2018</a>, 2018.
</mixed-citation></ref-html>
<ref-html id="bib1.bib17"><label>17</label><mixed-citation>
Elsig, J., Schmitt, J., Leuenberger, D., Schneider, R., Eyer, M.,
Leuenberger, M., Joos, F., Fischer, H., and Stocker, T. F.: Stable isotope
constraints on Holocene carbon cycle changes from an Antarctic ice core,
Nature, 461, 507–510, 2009.
</mixed-citation></ref-html>
<ref-html id="bib1.bib18"><label>18</label><mixed-citation>
Eyring, V., Bony, S., Meehl, G. A., Senior, C. A., Stevens, B., Stouffer, R.
J., and Taylor, K. E.: Overview of the Coupled Model Intercomparison Project
Phase 6 (CMIP6) experimental design and organization, Geosci. Model Dev., 9,
1937–1958, <a href="https://doi.org/10.5194/gmd-9-1937-2016" target="_blank">https://doi.org/10.5194/gmd-9-1937-2016</a>, 2016.
</mixed-citation></ref-html>
<ref-html id="bib1.bib19"><label>19</label><mixed-citation>
Fischer, N. and Jungclaus, J. H.: Evolution of the seasonal temperature
cycle in a transient Holocene simulation: orbital forcing and sea-ice, Clim.
Past, 7, 1139–1148, <a href="https://doi.org/10.5194/cp-7-1139-2011" target="_blank">https://doi.org/10.5194/cp-7-1139-2011</a>, 2011.
</mixed-citation></ref-html>
<ref-html id="bib1.bib20"><label>20</label><mixed-citation>
Ford, A. and Clarke, K. C.: Linking the past and present of the ancient Maya: lowland land use, population distribution, and density in the Late Classic Period, in: The Oxford Handbook of Historical Ecology and Applied Archaeology, edited by: Isendahl, C. and Stump, D., Oxford University Press, Oxford, <a href="https://doi.org/10.1093/oxfordhb/9780199672691.013.33" target="_blank">https://doi.org/10.1093/oxfordhb/9780199672691.013.33</a>, 2015.
</mixed-citation></ref-html>
<ref-html id="bib1.bib21"><label>21</label><mixed-citation>
Freeman, J., Baggio, J. A., Robinson, E., Byers, D. A., Gayo, E., Finley, J. B., Meyer, J. A., Kelly, R. L., and Anderies, J. M.: Synchronisation of energy consumption by human societies throughout the Holocene, P. Natl. Acad. Sci. USA, 115, 9962–9967, <a href="https://doi.org/10.1073/pnas.1802859115" target="_blank">https://doi.org/10.1073/pnas.1802859115</a>, 2018.
</mixed-citation></ref-html>
<ref-html id="bib1.bib22"><label>22</label><mixed-citation>
Fyfe, R. M., Woodbridge, J. E., and Roberts, N.: From forest to farmland:
pollen-inferred land cover change across Europe using the pseudobiomization
approach, Global Change Biol., 21: 1197–1212, <a href="https://doi.org/10.1111/gcb.12776" target="_blank">https://doi.org/10.1111/gcb.12776</a>, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib23"><label>23</label><mixed-citation>
Gaillard, M.-J.: First pollen-based REVEALS reconstruction of vegetation abundance in Europe for five time windows between 6&thinsp;k years BP and present, PANGAEA, <a href="https://doi.org/10.1594/PANGAEA.897303" target="_blank">https://doi.org/10.1594/PANGAEA.897303</a>, 2019.
</mixed-citation></ref-html>
<ref-html id="bib1.bib24"><label>24</label><mixed-citation>
Gaillard, M.-J., Sugita, S., Mazier, F., Kaplan, J. O., Trondman, A.-K., Brostroem, A., Hickler, T., Kjellstroem, E., Kunes, P., Lemmen, C., Olofsson, J., Smith, B., and Strandberg, G.: Holocene land-cover reconstructions for studies on land-cover feedbacks, Clim. Past, 6, 483–499,
<a href="https://doi.org/10.5194/cp-6-483-2010" target="_blank">https://doi.org/10.5194/cp-6-483-2010</a>, 2010.
</mixed-citation></ref-html>
<ref-html id="bib1.bib25"><label>25</label><mixed-citation>
Gaillard, M-J., Whitehouse, N., Madella, M., Morrison, K., and von Gunten, L.: Past land use and land cover, PAGES Mag.,26, 1–44,
<a href="https://doi.org/10.22498/pages.26.1" target="_blank">https://doi.org/10.22498/pages.26.1</a>, 2018.
</mixed-citation></ref-html>
<ref-html id="bib1.bib26"><label>26</label><mixed-citation>
Harrison, S. P.: Pollen-based reconstructions of bioclimatic variables at 6000&thinsp;yr&thinsp;BP, <a href="https://doi.org/10.5281/zenodo.3601028" target="_blank">https://doi.org/10.5281/zenodo.3601028</a>, 2020a.
</mixed-citation></ref-html>
<ref-html id="bib1.bib27"><label>27</label><mixed-citation>
Harrison, S. P.: Climate simulations of impact of LULC on mid-Holocene climate, <a href="https://doi.org/10.5281/zenodo.3601040" target="_blank">https://doi.org/10.5281/zenodo.3601040</a>, 2020b.
</mixed-citation></ref-html>
<ref-html id="bib1.bib28"><label>28</label><mixed-citation>
Harrison, S. P.: Code to compare model outputs and pollen-based reconstructions, <a href="https://doi.org/10.5281/zenodo.3601011" target="_blank">https://doi.org/10.5281/zenodo.3601011</a>, 2020c.
</mixed-citation></ref-html>
<ref-html id="bib1.bib29"><label>29</label><mixed-citation>
Harrison, S. P., Bartlein, P. J., Izumi, K., Li, G., Annan, J., Hargreaves,
J., Braconnot, P. B., and Kageyama, M.: Evaluation of CMIP5 palaeo-simulations to improve climate projections, Nat. Clim. Change, 5,
735–743, 2015.
</mixed-citation></ref-html>
<ref-html id="bib1.bib30"><label>30</label><mixed-citation>
He, F., Vavrus, S. J., Kutzbach, J. E., Ruddiman, W. F., Kaplan, J. O., and
Krumhardt, K. M.: Simulating global and local surface temperature changes due
to Holocene anthropogenic land cover change, Geophys. Res. Lett., 41, 623–631, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib31"><label>31</label><mixed-citation>
Hellman, S., Gaillard, M.-J., Broström, A., and Sugita, S.: The REVEALS
model, new tool to estimate past regional plant abundance from pollen data
in large lakes: validation in southern Sweden, J. Quatenary Sci., 22, 1–22,
2008a.
</mixed-citation></ref-html>
<ref-html id="bib1.bib32"><label>32</label><mixed-citation>
Hellman, S., Gaillard, M.-J., Broström, A., and Sugita, S.: Effects of
the sampling design and selection of parameter values on pollen-based
quantitative reconstructions of regional vegetation: a case study in southern Sweden using the REVEALS model, Veg. Hist. Archaeobot., 17, 445–460, 2008b.
</mixed-citation></ref-html>
<ref-html id="bib1.bib33"><label>33</label><mixed-citation>
Herrero, M., Havlík, P., Valin, H., Notenbaert, A., Rufino, M. C.,
Thornton, P. K., Blümmel, M., Weiss, F., Grace, D., and Obersteiner, M.:
Biomass use, production, feed efficiencies, and greenhouse gas emissions from global livestock systems, P. Natl. Acad. Sci. USA, 110, 20888–20893, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib34"><label>34</label><mixed-citation>
Hughes, R. E., Weiberg, E., Bonnier, A., Finne, M., and Kaplan, J. O.:
Quantifying land use in past societies from cultural practice and archaeological data, Land, 7, 9, <a href="https://doi.org/10.3390/land7010009" target="_blank">https://doi.org/10.3390/land7010009</a>, 2018.
</mixed-citation></ref-html>
<ref-html id="bib1.bib35"><label>35</label><mixed-citation>
Hurtt, G., Chini, L., Sahajpal, R., Frolking, S., Bodirsky, B. L., Calvin, K., Doelman, J., Fisk, J., Fujimori, S., Klein Goldewijk, K., Hasegawa, T., Havlik, P., Heinimann, A., Humpenöder, F., Jungclaus, J., Kaplan, J., Krisztin, T., Lawrence, D., Lawrence, P., Mertz, O., Pongratz, J., Popp, A., Riahi, K., Shevliakova, E., Stehfest, E., Thornton, P., van Vuuren, D., and Zhang, X.: Harmonization of global land use scenarios (LUH2): SSP585 v2.1f 2015–2100, Earth System Grid Federation, <a href="https://doi.org/10.22033/ESGF/input4MIPs.1662" target="_blank">https://doi.org/10.22033/ESGF/input4MIPs.1662</a>, 2017
</mixed-citation></ref-html>
<ref-html id="bib1.bib36"><label>36</label><mixed-citation>
Janssen, E., Poblome, J., Claeys, J., Kint, V., Degryse, P., Marinova, E., and Muys, B.: Fuel for debating ancient economies. Calculating wood consumption at urban scale in Roman Imperial times, J. Arch. Sci.: Rep., 11, 592–599, 2017.
</mixed-citation></ref-html>
<ref-html id="bib1.bib37"><label>37</label><mixed-citation>
Joos, F. and Spahni, R.: Rates of change in natural and anthropogenic radiative forcing over the past 20,000 years, P. Natl. Acad. Sci. USA, 105,
1425–1430, 2008.
</mixed-citation></ref-html>
<ref-html id="bib1.bib38"><label>38</label><mixed-citation>
Joos, F., Gerber, S., Prentice, I. C., Otto-Bliesner, B. L., and Valdes, P. J.: Transient simulations of Holocene atmospheric carbon dioxide and terrestrial carbon since the last glacial maximum, Global Biogeochem. Cy., 18, GB2002, <a href="https://doi.org/10.1029/2003GB002156" target="_blank">https://doi.org/10.1029/2003GB002156</a>, 2004.
</mixed-citation></ref-html>
<ref-html id="bib1.bib39"><label>39</label><mixed-citation>
Jungclaus, J. H., Bard, E., Baroni, M., Braconnot, P., Cao, J., Chini, L. P.,
Egorova, T., Evans, M., González-Rouco, J. F., Goosse, H., Hurtt, G. C.,
Joos, F., Kaplan, J. O., Khodri, M., Klein Goldewijk, K., Krivova, N.,
LeGrande, A. N., Lorenz, S. J., Luterbacher, J., Man, W., Maycock, A. C.,
Meinshausen, M., Moberg, A., Muscheler, R., Nehrbass-Ahles, C.,
Otto-Bliesner, B. I., Phipps, S. J., Pongratz, J., Rozanov, E., Schmidt, G. A., Schmidt, H., Schmutz, W., Schurer, A., Shapiro, A. I., Sigl, M., Smerdon,
J. E., Solanki, S. K., Timmreck, C., Toohey, M., Usoskin, I. G., Wagner, S.,
Wu, C.-J., Yeo, K. L., Zanchettin, D., Zhang, Q., and Zorita, E.: The PMIP4
contribution to CMIP6 – Part 3: The last millennium, scientific objective,
and experimental design for the PMIP4 past1000 simulations, Geosci. Model
Dev., 10, 4005–4033, <a href="https://doi.org/10.5194/gmd-10-4005-2017" target="_blank">https://doi.org/10.5194/gmd-10-4005-2017</a>, 2017.
</mixed-citation></ref-html>
<ref-html id="bib1.bib40"><label>40</label><mixed-citation>
Kageyama, M., Braconnot, P., Harrison, S. P., Haywood, A., Jungclaus, J.,
Otto-Bliesner, B., Peterschmitt, J.-Y., Abe-Ouchi, A., Albani, S., Bartlein,
P., Brierley, C., Crucifix, M., Dolan, A., Fernandez-Donado, L., Fischer, H., Hopcroft, P., Ivanovic, R., Lambert, F., Lunt, D., Mahowald, N., Peltier, W. R., Phipps, S., Roche, D., Schmidt, G., Tarasov, L., Valdes, P., Zhang, Q., and Zhou, T.: The PMIP4 contribution to CMIP6 – Part 1: Overview and over-arching analysis plan, Geosci. Model Dev., 11, 1033–1057,
<a href="https://doi.org/10.5194/gmd-11-1033-2018" target="_blank">https://doi.org/10.5194/gmd-11-1033-2018</a>, 2018.
</mixed-citation></ref-html>
<ref-html id="bib1.bib41"><label>41</label><mixed-citation>
Kaplan, J. O.: ALCC_Comparison_Figure, available at: <a href="https://github.com/jedokaplan/ALCC_comparison_figure" target="_blank"/>, last access: 14 February 2020.
</mixed-citation></ref-html>
<ref-html id="bib1.bib42"><label>42</label><mixed-citation>
Kaplan, J. O. and Krumhardt, K. M.: The KK10 Anthropogenic Land Cover Change scenario for the preindustrial Holocene, PANGAEA, <a href="https://doi.org/10.1594/PANGAEA.871369" target="_blank">https://doi.org/10.1594/PANGAEA.871369</a>, 2011.
</mixed-citation></ref-html>
<ref-html id="bib1.bib43"><label>43</label><mixed-citation>
Kaplan, J. O., Krumhardt, K. M., Ellis, E. C., Ruddiman, W. F., Lemmen, C., and Klein Goldewijk, K.: Holocene carbon emissions as a result of anthropogenic land cover change, Holocene, 21, 775–791, 2011.
</mixed-citation></ref-html>
<ref-html id="bib1.bib44"><label>44</label><mixed-citation>
Kaplan, J. O., Krumhardt, K. M., Gaillard, M.-J., Sugita, S., Trondman, A.-K., Fyfe, R., Marquer, L., Mazier, F., and Nielsen, A. B.: Constraining the deforestation history of Europe: Evaluation of historical land use scenarios with pollen-based land cover reconstructions, Land, 6, 9, <a href="https://doi.org/10.3390/land6040091" target="_blank">https://doi.org/10.3390/land6040091</a>, 2017.
</mixed-citation></ref-html>
<ref-html id="bib1.bib45"><label>45</label><mixed-citation>
Kaufman, D., McKay, N., Routson, C., Erb, M., Davis, B., Heiri, O., Jaccard,
S., Tierney, J., Dätwyler, C., et al.: A global database of Holocene paleo-temperature records, Scient. Data, in press, 2020.
</mixed-citation></ref-html>
<ref-html id="bib1.bib46"><label>46</label><mixed-citation>
Kay, A. U., Fuller, D. Q., Neumann, K., Eichhorn, B., Höhn, A., Morin-Rivat, J., Champion, L., Linseele, V., Huysecom, E., Ozainne, S., Lespez, L., Biagetti, S., Madella, M., Salzmann, U., and Kaplan, J. O.:
Diversification, intensification, and specialization: Changing land use in
western Africa from 1800&thinsp;BC to AD&thinsp;1500, J. World Prehist., 32, 179–228,
<a href="https://doi.org/10.1007/s10963-019-09131-2" target="_blank">https://doi.org/10.1007/s10963-019-09131-2</a>, 2019.
</mixed-citation></ref-html>
<ref-html id="bib1.bib47"><label>47</label><mixed-citation>
Klein Goldewijk, C. G. M.: Anthropogenic land-use estimates for the Holocene; HYDE 3.2. DANS, <a href="https://doi.org/10.17026/dans-25g-gez3" target="_blank">https://doi.org/10.17026/dans-25g-gez3</a>, 2017.
</mixed-citation></ref-html>
<ref-html id="bib1.bib48"><label>48</label><mixed-citation>
Klein Goldewijk, K., Beusen, A., van Drecht, G., and de Vos, M.: The HYDE 3.1 spatially explicit database of human induced land use change over the past 12,000 years, Global Ecol. Biogeogr., 20, 73–86, 2011.
</mixed-citation></ref-html>
<ref-html id="bib1.bib49"><label>49</label><mixed-citation>
Klein Goldewijk, K., Beusen, A., Doelman, J., and Stehfest, E.: Anthropogenic land-use estimates for the Holocene; HYDE 3.2, Earth Syst. Sci. Data, 9, 927–953, <a href="https://doi.org/10.5194/essd-9-927-2017" target="_blank">https://doi.org/10.5194/essd-9-927-2017</a>, 2017a.
</mixed-citation></ref-html>
<ref-html id="bib1.bib50"><label>50</label><mixed-citation>
Klein Goldewijk, K., Dekker, S. C., and van Zanden, J. L.: Per-capita
estimations of long-term historical land use and the consequences for global
change research, J. Land Use Sci., 12, 313–337, <a href="https://doi.org/10.1080/1747423X.2017.1354938" target="_blank">https://doi.org/10.1080/1747423X.2017.1354938</a>, 2017b.
</mixed-citation></ref-html>
<ref-html id="bib1.bib51"><label>51</label><mixed-citation>
Lawrence, D. M., Hurtt, G. C., Arneth, A., Brovkin, V., Calvin, K. V., Jones,
A. D., Jones, C. D., Lawrence, P. J., de Noblet-Ducoudré, N., Pongratz, J., Seneviratne, S. I., and Shevliakova, E.: The Land Use Model Intercomparison Project (LUMIP) contribution to CMIP6: rationale and experimental design, Geosci. Model Dev., 9, 2973–2998, <a href="https://doi.org/10.5194/gmd-9-2973-2016" target="_blank">https://doi.org/10.5194/gmd-9-2973-2016</a>, 2016.
</mixed-citation></ref-html>
<ref-html id="bib1.bib52"><label>52</label><mixed-citation>
Le Quéré, C., Andrew, R. M., Friedlingstein, P., Sitch, S., Hauck, J., Pongratz, J., Pickers, P. A., Korsbakken, J. I., Peters, G. P., Canadell,
J. G., Arneth, A., Arora, V. K., Barbero, L., Bastos, A., Bopp, L., Chevallier, F., Chini, L. P., Ciais, P., Doney, S. C., Gkritzalis, T., Goll,
D. S., Harris, I., Haverd, V., Hoffman, F. M., Hoppema, M., Houghton, R. A.,
Hurtt, G., Ilyina, T., Jain, A. K., Johannessen, T., Jones, C. D., Kato, E.,
Keeling, R. F., Goldewijk, K. K., Landschützer, P., Lefèvre, N., Lienert, S., Liu, Z., Lombardozzi, D., Metzl, N., Munro, D .R., Nabel, J. E. M. S., Nakaoka, S.-I., Neill, C., Olsen, A., Ono, T., Patra, P., Peregon, A., Peters, W., Peylin, P., Pfeil, B., Pierrot, D., Poulter, B., Rehder, G.,
Resplandy, L., Robertson, E., Rocher, M., Rödenbeck, C., Schuster, U.,
Schwinger, J., Séférian, R., Skjelvan, I., Steinhoff, T., Sutton, A., Tans, P. P., Tian, H., Tilbrook, B., Tubiello, F. N., van der Laan-Luijkx, I. T., van der Werf, G. R., Viovy, N., Walker, A. P., Wiltshire, A. J., Wright, R., Zaehle, S., and Zheng, B.: Global Carbon Budget 2018, Earth Syst. Sci. Data, 10, 2141–2194, <a href="https://doi.org/10.5194/essd-10-2141-2018" target="_blank">https://doi.org/10.5194/essd-10-2141-2018</a>, 2018.
</mixed-citation></ref-html>
<ref-html id="bib1.bib53"><label>53</label><mixed-citation>
Lewin, S. and Whitehouse, N.: Regional Land Use mapping code and dataset for Early and Middle Neolithic Ireland (3750–3600&thinsp;cal&thinsp;BC and 3600–3400&thinsp;cal&thinsp;BC, respectively), <a href="https://doi.org/10.5281/zenodo.3625226" target="_blank">https://doi.org/10.5281/zenodo.3625226</a>, 2020.
</mixed-citation></ref-html>
<ref-html id="bib1.bib54"><label>54</label><mixed-citation>
Loisel, J., Yu, Z., Beilman, D. W., Camill, P., Alm, J., Amesbury, M. J.,
Anderson, D., Andersson, S., Bochicchio, C., Barber, K., Belyea, L. R., Bunbury, J., Chambers, F. M., Charman, D. J., Vleeschouwer, F.D., Fiałkiewicz-Kozieł, B., Finkelstein, S. A., Gałka, M., Garneau, M.,
Hammarlund, D., Hinchcliffe, W., Holmquist, J., Hughes, P., Jones, M. C.,
Klein, E. S., Kokfelt, U., Korhola, A., Kuhry, P., Lamarre, A., Lamentowicz,
M., Large, D., Lavoie, M., MacDonald, G., Magnan, G., Mäkilä, M.,
Mallon, G., Mathijssen, P., Mauquoy, D., McCarroll, J., Moore, T. R., Nichols, J., O'Reilly, B., Oksanen, P., Packalen, M., Peteet, D., Richard,
P. J., Robinson, S., Ronkainen, T., Rundgren, M., Sannel, A. B. K., Tarnocai,
C., Thom, T., Tuittila, E.-S., Turetsky, M., Väliranta, M., and der Linden, M., van Geel B., van Bellen, S., Vitt, D., Zhao, Y., and Zhou, W.: A
database and synthesis of northern peatland soil properties and Holocene carbon and nitrogen accumulation, Holocene, 24, 1028–1042, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib55"><label>55</label><mixed-citation>
Lyman, R. L.: Quantitative Paleozoology, Cambridge University Press, Cambridge, 2008.
</mixed-citation></ref-html>
<ref-html id="bib1.bib56"><label>56</label><mixed-citation>
Maezumi, S. Y., Robinson, M., de Souza, J., Urrego, D. H., Schaan, D., Alves,
D., and Iriarte, J.: New insights from pre-Columbian land use and fire management in Amazonian Dark Earth forests, Front. Ecol. Evol., 6, 111, <a href="https://doi.org/10.3389/fevo.2018.00111" target="_blank">https://doi.org/10.3389/fevo.2018.00111</a>, 2018.
</mixed-citation></ref-html>
<ref-html id="bib1.bib57"><label>57</label><mixed-citation>
Mahowald, N. M., Randerson, J. T., Lindsay, K., Munoz, E., Doney, S. C.,
Lawrence, P., Schlunegger, S., Ward, D. S., Lawrence, D., and Hoffman, F. M.: Interactions between land use change and carbon cycle feedbacks, Global
Biogeochem. Cy., 31, 96–113, <a href="https://doi.org/10.1002/2016GB005374" target="_blank">https://doi.org/10.1002/2016GB005374</a>, 2017.
</mixed-citation></ref-html>
<ref-html id="bib1.bib58"><label>58</label><mixed-citation>
Marston, J. M.: Modeling wood aquisition strategies from archaeological charcoal remains, J. Arch. Sci., 36, 2192–2200, 2009.
</mixed-citation></ref-html>
<ref-html id="bib1.bib59"><label>59</label><mixed-citation>
Marston, J. M., Holdaway, S. J., and Wendrich, W.: Early- and middle-Holocene
wood exploitation in the Fayum basin, Egypt, Holocene, 27, 1812–1824, 2017.
</mixed-citation></ref-html>
<ref-html id="bib1.bib60"><label>60</label><mixed-citation>
Mauri, A., Davis, B. A. S., Collins, P. M., and Kaplan, J. O.: The influence
of atmospheric circulation on the mid-Holocene climate of Europe: a data–model comparison, Clim. Past, 10, 1925–1938,
<a href="https://doi.org/10.5194/cp-10-1925-2014" target="_blank">https://doi.org/10.5194/cp-10-1925-2014</a>, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib61"><label>61</label><mixed-citation>
Mazoyer, M. and Roudart, L.: A History of World Agriculture: From the Neolithic to the Current Crisis, Earthscan, UK, 2006.
</mixed-citation></ref-html>
<ref-html id="bib1.bib62"><label>62</label><mixed-citation>
McGrath, M. J., Luyssaert, S., Meyfroidt, P., Kaplan, J. O., Burgi, M., Chen, Y., Erb, K., Gimmi, U., McInerney, D., Naudts, K., Otto, J., Pasztor, F., Ryder, J., Schelhaas, M. J., and Valade, A.: Reconstructing European forest management from 1600 to 2010, Biogeoscience, 12, 4291–4316,
<a href="https://doi.org/10.5194/bg-12-4291-2015" target="_blank">https://doi.org/10.5194/bg-12-4291-2015</a>, 2015.
</mixed-citation></ref-html>
<ref-html id="bib1.bib63"><label>63</label><mixed-citation>
McLaughlin, T. R., Whitehouse, N. J., Schulting, R. J., McClatchie, M., Barratt, P., and Bogaard, A.: The changing face of Neolithic and Bronze Age
Ireland: A big data approach to the settlement and burial records, J. World
Prehist., 29, 117–153, <a href="https://doi.org/10.1007/s10963-016-9093-0" target="_blank">https://doi.org/10.1007/s10963-016-9093-0</a>, 2016.
</mixed-citation></ref-html>
<ref-html id="bib1.bib64"><label>64</label><mixed-citation>
Messori, G., Gaetani, M., Zhang, Q., Zhang, Q., and Pausata, F. S. R.: The
water cycle of the mid-Holocene West African monsoon: The role of vegetation
and dust emission changes, Int. J. Climatol., 39, 1927–1939, <a href="https://doi.org/10.1002/joc.5924" target="_blank">https://doi.org/10.1002/joc.5924</a>, 2019.
</mixed-citation></ref-html>
<ref-html id="bib1.bib65"><label>65</label><mixed-citation>
Mitchell, L., Brook, E., Lee, J., Buizert, C., and Sowers, T.: Constraints on the late Holocene anthropogenic contribution to the atmospheric methane budget, Science, 342, 964–966, <a href="https://doi.org/10.1126/science.1238920" target="_blank">https://doi.org/10.1126/science.1238920</a>, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib66"><label>66</label><mixed-citation>
Morrison, K. D., Hammer, E., Popova, L., Madella, M., Whitehouse, N., Gaillard, M.-J., and LandCover6k Land-Use Group Members: Global-scale
comparisons of human land use: developing shared terminology for land-use
practices for global changes, PAGES Mag., 26, 8–9, 2018.
</mixed-citation></ref-html>
<ref-html id="bib1.bib67"><label>67</label><mixed-citation>
Myhre, G., Shindell, D., Bréon, F.-M., Collins, W., Fuglestvedt, J., Huang, J., Koch, D., Lamarque, J.-F., Lee, D., Mendoza, B., Nakajima, T.,
Robock, A., Stephens, G., Takemura T., and Zhang, H.: Anthropogenic and
natural radiative forcing, in: Climate Change 2013: The Physical Science
Basis. Contribution of Working Group I to the Fifth Assessment Report of the
Intergovernmental Panel on Climate Change, edited by: Stocker, T. F., Qin, D., Plattner, G.-K., Tignor, M., Allen, S. K., Boschung, J., Nauels, A., Xia, Y., Bex, V., and Midgley, P. M., Cambridge University Press Cambridge, Cambridge, UK and New York, NY, USA, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib68"><label>68</label><mixed-citation>
Nakagawa, T., Tarasov, P. E., Nishida, K., Gotanda, K., and Yasuda, Y.:
Quantitative pollen-based climate reconstruction in central Japan: application to surface and Late Quaternary spectra, Quaternary Sci. Rev., 21,
2099–2113, 2002.
</mixed-citation></ref-html>
<ref-html id="bib1.bib69"><label>69</label><mixed-citation>
Nielsen, A. B., Giesecke, T., Theuerkauf, M., Feeser, I., Behre, K.-H., Beug,
H.-J., Chen, S.-H., Christiansen, J., Dörfler, W., Endtmann, E., Jahns, S., de Klerk, O., Kühl, N., Latałowa, M., Odgaard, B. V., Rasmussen,
P., Stockholm, J. R., Voigt, R., Wiethold, J., and Wolters, S.: Quantitative
reconstructions of changes in regional openness in north-central Europe
reveal new insights into old questions, Quaternary Sci. Rev., 47, 131–149, 2012.
</mixed-citation></ref-html>
<ref-html id="bib1.bib70"><label>70</label><mixed-citation>
Oh, Y., Conte, M., Kang, S., Kim, J., and Hwang, J.: Population fluctuation
and the adoption of food production in prehistoric Korea: using radiocarbon
dates as a proxy for population change, Radiocarbon, 59, 1761–1770, <a href="https://doi.org/10.1017/RDC.2017.122" target="_blank">https://doi.org/10.1017/RDC.2017.122</a>, 2017.
</mixed-citation></ref-html>
<ref-html id="bib1.bib71"><label>71</label><mixed-citation>
Ortega-Rosas, C. I., Guiot, J., Penalba, M. C., and Ortiz-Acosta, M. E.:
Biomization and quantitative climate reconstruction techniques in northwestern Mexico – with an application to four Holocene pollen sequences,
Global Planet. Change, 61, 242–266, 2008.
</mixed-citation></ref-html>
<ref-html id="bib1.bib72"><label>72</label><mixed-citation>
Orton, D., Gaastra, J., and Vander Linden, M.: Between the Danube and the Deep Blue Sea: zooarchaeological meta-analysis reveals variability in the
spread and development of Neolithic farming across the western Balkans, Open
Quat., 2, 6, <a href="https://doi.org/10.5334/oq.28" target="_blank">https://doi.org/10.5334/oq.28</a>, 2016.
</mixed-citation></ref-html>
<ref-html id="bib1.bib73"><label>73</label><mixed-citation>
Otto-Bliesner, B. L., Braconnot, P., Harrison, S. P., Lunt, D. J., Abe-Ouchi, A., Albani, S., Bartlein, P. J., Capron, E., Carlson, A. E., Dutton, A., Fischer, H., Goelzer, H., Govin, A., Haywood, A., Joos, F., LeGrande, A. N., Lipscomb, W. H., Lohmann, G., Mahowald, N., Nehrbass-Ahles, C., Pausata, F. S. R., Peterschmitt, J.-Y., Phipps, S. J., Renssen, H., and Zhang, Q.: The PMIP4 contribution to CMIP6 – Part 2: Two interglacials, scientific objective and experimental design for Holocene and Last Interglacial simulations, Geosci. Model Dev., 10, 3979–4003, <a href="https://doi.org/10.5194/gmd-10-3979-2017" target="_blank">https://doi.org/10.5194/gmd-10-3979-2017</a>, 2017.
</mixed-citation></ref-html>
<ref-html id="bib1.bib74"><label>74</label><mixed-citation>
Pausata, F. S. R., Messori, G., and Zhang, Q.: Impacts of dust reduction on the northward expansion of the African monsoon during the Green Sahara period, Earth Planet. Sc. Lett., 434, 298–307, <a href="https://doi.org/10.1016/j.epsl.2015.11.049" target="_blank">https://doi.org/10.1016/j.epsl.2015.11.049</a>, 2016.
</mixed-citation></ref-html>
<ref-html id="bib1.bib75"><label>75</label><mixed-citation>
Perugini, L. Caporaso, L., Marconi, S., Cescatti, A., Quesada, B., de Noblet-Ducoudré, N., House, J. I., and Arneth, A.: Biophysical effects on temperature and precipitation due to land cover change, Environ. Res. Lett., 12, 053002, <a href="https://doi.org/10.1088/1748-9326/aa6b3f" target="_blank">https://doi.org/10.1088/1748-9326/aa6b3f</a>, 2017.
</mixed-citation></ref-html>
<ref-html id="bib1.bib76"><label>76</label><mixed-citation>
Phelps, L. N. and Kaplan, J. O.: Land use for animal production in global
change studies: Defining and characterizing a framework, Global Change Biol.,
23, 4457–4471, <a href="https://doi.org/10.1111/gcb.13732" target="_blank">https://doi.org/10.1111/gcb.13732</a>, 2017.
</mixed-citation></ref-html>
<ref-html id="bib1.bib77"><label>77</label><mixed-citation>
Pirzamanbein, B., Lindström, J., Poska, A., Sugita, S., Trondman, A.,
Fyfe, R., Mazier, F., Nielsen, A. B., Kaplan, J. O., Bjune, A. E., Birks, H. J. B., Giesecke, T., Kangur, M., Latałowa, M., Marquer, L., Smith, B., and Gaillard, M.-J.: Creating spatially continuous maps of past land cover from point estimates: A new statistical approach applied to pollen data, Ecol. Complex., 20, 127–141, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib78"><label>78</label><mixed-citation>
Pirzamanbein, B., Lindström, J., Poska, A., and Gaillard, M.-J.: Modelling spatial compositional data: Reconstructions of past land cover and
uncertainties, Spat. Stat., 24, 14–31, 2018.
</mixed-citation></ref-html>
<ref-html id="bib1.bib79"><label>79</label><mixed-citation>
Pongratz, J., Reick, C., Raddatz, T., and Claussen, M.: A reconstruction of
global agricultural areas and land cover for the last millennium, Global
Biogeochem. Cy., 22, GB3018, <a href="https://doi.org/10.1029/2007GB003153" target="_blank">https://doi.org/10.1029/2007GB003153</a>, 2008.
</mixed-citation></ref-html>
<ref-html id="bib1.bib80"><label>80</label><mixed-citation>
Pongratz, J., Reick, C. H., Raddatz, T., and Claussen, M.: Biogeophysical
versus biogeochemical climate response to historical anthropogenic land cover change, Geophys. Res. Lett., 37, L08702, <a href="https://doi.org/10.1029/2010GL043010" target="_blank">https://doi.org/10.1029/2010GL043010</a>, 2010.
</mixed-citation></ref-html>
<ref-html id="bib1.bib81"><label>81</label><mixed-citation>
Pongratz, J., Reick, C. H., Houghton, R. A., and House, J. I.: Terminology as a key uncertainty in net land use and land cover change carbon flux estimates, Earth Syst. Dynam., 5, 177–195, <a href="https://doi.org/10.5194/esd-5-177-2014" target="_blank">https://doi.org/10.5194/esd-5-177-2014</a>, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib82"><label>82</label><mixed-citation>
Ramankutty, N. and Foley, J. A.: Estimating historical changes in global land cover: Croplands from 1700 to 1992, Global Biogeochem. Cy., 13, 997–1027, 1999.
</mixed-citation></ref-html>
<ref-html id="bib1.bib83"><label>83</label><mixed-citation>
Reitz, E. J. and Wing, E. S.: Zooarchaeology, Cambridge University Press,
Cambridge, 2008.
</mixed-citation></ref-html>
<ref-html id="bib1.bib84"><label>84</label><mixed-citation>
Rick, J. W.: Dates as data: an examination of the Peruvian Preceramic
radiocarbon record, Am. Antiq., 52, 55–73, 1987.
</mixed-citation></ref-html>
<ref-html id="bib1.bib85"><label>85</label><mixed-citation>
Robinson, E., Zahid, H. J., Codding, B. F., Haas, R., and Kelly, R. L.:
Spatiotemporal dynamics of prehistoric human population growth: radiocarbon
`dates as data' and population ecology models, J. Arch. Sci., 101, 63–71,
2019.
</mixed-citation></ref-html>
<ref-html id="bib1.bib86"><label>86</label><mixed-citation>
Ruddiman, W. F.: The anthropogenic greenhouse era began thousands of years ago, Climatic Change, 61, 261–293, <a href="https://doi.org/10.1023/B:CLIM.0000004577.17928.fa" target="_blank">https://doi.org/10.1023/B:CLIM.0000004577.17928.fa</a>,
2003.
</mixed-citation></ref-html>
<ref-html id="bib1.bib87"><label>87</label><mixed-citation>
Russell, T., Silva, F., and Steele, J.: Modelling the spread of farming in
the Bantu-speaking regions of Africa: an archaeology-based phylogeography,
Plos One, 9, e87584, <a href="https://doi.org/10.1371/journal.pone.0087854" target="_blank">https://doi.org/10.1371/journal.pone.0087854</a>, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib88"><label>88</label><mixed-citation>
Shennan, S., Downey, S. S., Timpson, A., Edinborough, K., Colledge, S., Kerig, T., Manning, K., and Thomas, M. G.: Regional population collapse followed initial agriculture booms in mid-Holocene Europe, Nat. Commun., 4,
248, <a href="https://doi.org/10.1038/ncomms3486" target="_blank">https://doi.org/10.1038/ncomms3486</a>, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib89"><label>89</label><mixed-citation>
Shevliakova, E., Pacala, S. W., Malyshev, S., Hurtt, G. C., Milly, P. C. D.,
Caspersen, J. P., Sentman, L. T., Fisk, J. P., Wirth, C., and Crevoisier, C.: Carbon cycling under 300 years of land use change: Importance of the secondary vegetation sink, Global Biogeochem. Cy., 23, GB2022,
<a href="https://doi.org/10.1029/2007GB003176" target="_blank">https://doi.org/10.1029/2007GB003176</a>, 2009.
</mixed-citation></ref-html>
<ref-html id="bib1.bib90"><label>90</label><mixed-citation>
Sigl, M., Winstrup, M., McConnell, J. R., Welten, K. C., Plunkett, G., Ludlow, F., Büntgen, U., Caffee, M., Chellman, N., Dahl-Jensen, D., Fischer, H., Kipfstuhl, S., Kostick, C., Maselli, O. J., Mekhaldi, F.,
Mulvaney, R., Muscheler, R., Pasteris, D. R., Pilcher, J. R., Salzer, M.,
Schüpbach, S., Steffensen, J. P., Vinther, B. M., and Woodruff, T. E.: Timing and climate forcing of volcanic eruptions for the past 2,500 years, Nature, 523, 543–549, <a href="https://doi.org/10.1038/nature14565" target="_blank">https://doi.org/10.1038/nature14565</a>, 2015.
</mixed-citation></ref-html>
<ref-html id="bib1.bib91"><label>91</label><mixed-citation>
Silva, F. and Steele, J.: New methods for reconstructing geographical effects on dispersal rates from large-scale radiocarbon databases, J. Arch. Sci., 52, 609–620, <a href="https://doi.org/10.1016/j.jas.2014.04.021" target="_blank">https://doi.org/10.1016/j.jas.2014.04.021</a>, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib92"><label>92</label><mixed-citation>
Silva, F. and Vander Linden, M.: Amplitude of travelling front as inferred
from <sup>14</sup>C predicts levels of genetic admixture among European early
farmers, Sci. Rep., 7, 11985, <a href="https://doi.org/10.1038/s41598-017-12318-2" target="_blank">https://doi.org/10.1038/s41598-017-12318-2</a>, 2017.
</mixed-citation></ref-html>
<ref-html id="bib1.bib93"><label>93</label><mixed-citation>
Silva, F., Stevens, C. J., Weisskopf, A., Castillo, C., Qin, L., Bevan, A.,
and Fuller, D. Q.: Modelling the geographical origin of rice cultivation in
Asia using the Rice Archaeological Database, Plos One, 10, e0137024, <a href="https://doi.org/10.1371/journal.pone.0137024" target="_blank">https://doi.org/10.1371/journal.pone.0137024</a>, 2015.
</mixed-citation></ref-html>
<ref-html id="bib1.bib94"><label>94</label><mixed-citation>
Singarayer, J. S., Valdes, P. J., Friedlingstein, P., Nelson, S., and Beerling, D. J.: Late Holocene methane rise caused by orbitally controlled
increase in tropical sources, Nature, 470, 82–85, <a href="https://doi.org/10.1038/nature09739" target="_blank">https://doi.org/10.1038/nature09739</a>,
2011.
</mixed-citation></ref-html>
<ref-html id="bib1.bib95"><label>95</label><mixed-citation>
Smith, M. C., Singarayer, J. S., Valdes, P. J., Kaplan, J. O., and Branch, N. P.: The biogeophysical climatic impacts of anthropogenic land use change during the Holocene, Clim. Past, 12, 923–941, <a href="https://doi.org/10.5194/cp-12-923-2016" target="_blank">https://doi.org/10.5194/cp-12-923-2016</a>, 2016.
</mixed-citation></ref-html>
<ref-html id="bib1.bib96"><label>96</label><mixed-citation>
Steinhilber, F., Abreu, J. A., Beer, J., Brunner, I., Christl, M., Fischer,
H., Heikkilä, U., Kubik, P. W., Mann, M., McCracken, K. G., Miller, H.,
Miyahara, H., Oerter, H., and Wilhelms, F.: 9400 years of cosmic radiation
and solar activity from ice cores and tree rings, P. Natl. Acad. Sci. USA, 109, 5967–5971, 2012.
</mixed-citation></ref-html>
<ref-html id="bib1.bib97"><label>97</label><mixed-citation>
Stocker, B. D., Strassmann, K., and Joos, F.: Sensitivity of Holocene atmospheric CO<sub>2</sub> and the modern carbon budget to early human land use: analyses with a process-based model, Biogeosciences, 8, 69–88,
<a href="https://doi.org/10.5194/bg-8-69-2011" target="_blank">https://doi.org/10.5194/bg-8-69-2011</a>, 2011.
</mixed-citation></ref-html>
<ref-html id="bib1.bib98"><label>98</label><mixed-citation>
Stocker, B. D., Yu, Z., Massa, C., and Joos, F.: Holocene peatland and
ice-core data constraints on the timing and magnitude of CO<sub>2</sub> emissions from past land use, P. Natl. Acad. Sci. USA, 114, 1492–1497,
<a href="https://doi.org/10.1073/pnas.1613889114" target="_blank">https://doi.org/10.1073/pnas.1613889114</a>, 2017.
</mixed-citation></ref-html>
<ref-html id="bib1.bib99"><label>99</label><mixed-citation>
Styring, A., Rösch, M., Stephan, E., Stika, H.-P., Fischer, E., Sillmann, E., and Bogaard, A.: Centralisation and long-term change in farming regimes: comparing agricultural practices in Neolithic and Iron Age south-west Germany, Proc. Prehist. Soc., 83, 357–381, <a href="https://doi.org/10.1017/ppr.2017.3" target="_blank">https://doi.org/10.1017/ppr.2017.3</a>, 2017.
</mixed-citation></ref-html>
<ref-html id="bib1.bib100"><label>100</label><mixed-citation>
Sugita, S.: Theory of quantitative reconstruction of vegetation I: pollen from large sites REVEALS regional vegetation composition, Holocene, 17,
229–241, 2007.
</mixed-citation></ref-html>
<ref-html id="bib1.bib101"><label>101</label><mixed-citation>
Tarasov, P., Williams, J. W., Andreev, A., Nakagawa, T., Bezrukova, E.,
Herzschuh, U., Igarashi, Y., Müller, S., Werner, K., and Zheng, Z.:
Satellite- and pollen-based quantitative woody cover reconstructions for
northern Asia: Verification and application to late-Quaternary pollen data,
Earth Planet. Sc. Lett., 264, 284–298, 2007.
</mixed-citation></ref-html>
<ref-html id="bib1.bib102"><label>102</label><mixed-citation>
Tauger, M. B.: Agriculture in World History, Routledge, Oxford, 2013.
</mixed-citation></ref-html>
<ref-html id="bib1.bib103"><label>103</label><mixed-citation>
Tierney, J. E., Pausata, F. S. R., and deMenocal, P. B.: Rainfall regimes of the Green Sahara, Sci. Adv., 3, e1601503, <a href="https://doi.org/10.1126/sciadv.1601503" target="_blank">https://doi.org/10.1126/sciadv.1601503</a>, 2017.
</mixed-citation></ref-html>
<ref-html id="bib1.bib104"><label>104</label><mixed-citation>
Timpson, A., Colledge, S., Crema, E., Edinborough, K., Kerig, T., Manning, K., Thomas, M. G., and Shennan, S.: Reconstructing regional population
fluctuations in the European Neolithic using radiocarbon dates: a new case-study using an improved method, J. Arch. Sci., 52, 549–557,
<a href="https://doi.org/10.1016/j.jas.2014.08.011" target="_blank">https://doi.org/10.1016/j.jas.2014.08.011</a>, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib105"><label>105</label><mixed-citation>
Toohey, M. and Sigl, M.: Volcanic stratospheric sulphur injections and
aerosol optical depth from 500&thinsp;BCE to 1900&thinsp;CE, Earth Syst. Sci. Data, 9,
809–831, <a href="https://doi.org/10.5194/essd-9-809-2017" target="_blank">https://doi.org/10.5194/essd-9-809-2017</a>, 2017.
</mixed-citation></ref-html>
<ref-html id="bib1.bib106"><label>106</label><mixed-citation>
Trondman, A. K., Gaillard, M.-J., Mazier, F., Sugita, S., Fyfe, R., Nielsen,
A. B., Twiddle, C., Barratt, P., Birks, H. J. B., Bjune, A. E., Björkman,
L., Broström, A., Caseldine, C., David, R., Dodson, J., Dörfler, W.,
Fischer, E., van Geel, B., Giesecke, T., Hultberg, T., Kalnina, L., Kangur,
M., van der Knaap, P., Koff, T., Kuneš, P., Lagerås, P., Latałowa,
M., Lechterbeck, J., Leroyer, C., Leydet, M., Lindbladh, M., Marquer, L.,
Mitchell, F. J. G., Odgaard, B. V., Peglar, S. M., Persson, T., Poska, A.,
Rösch, M., Seppä, H., Veski, S., and Wick, L.: Pollen-based quantitative reconstructions of Holocene regional vegetation cover (plant-functional types and land-cover types) in Europe suitable for climate
modelling, Global Change Biol., 21, 676–697, <a href="https://doi.org/10.1111/gcb.12737" target="_blank">https://doi.org/10.1111/gcb.12737</a>, 2015.
</mixed-citation></ref-html>
<ref-html id="bib1.bib107"><label>107</label><mixed-citation>
Trondman, A.-K., Gaillard, M.-J., Sugita, S., Björkman, L., Greisman, A., Hultberg, T., Lagerås, P., and Lindbladh, M.: Are pollen records from small sites appropriate for REVEALS model-based quantitative reconstructions of past regional vegetation? An empirical test in southern Sweden, Veget. Hist. Archaeobot., 25, 131–151, <a href="https://doi.org/10.1007/s00334-015-0536-9" target="_blank">https://doi.org/10.1007/s00334-015-0536-9</a>, 2016.
</mixed-citation></ref-html>
<ref-html id="bib1.bib108"><label>108</label><mixed-citation>
van den Hurk, B., Kim, H., Krinner, G., Seneviratne, S. I., Derksen, C., Oki,
T., Douville, H., Colin, J., Ducharne, A., Cheruy, F., Viovy, N., Puma, M. J., Wada, Y., Li, W., Jia, B., Alessandri, A., Lawrence, D. M., Weedon, G. P., Ellis, R., Hagemann, S., Mao, J., Flanner, M. G., Zampieri, M., Materia, S., Law, R. M., and Sheffield, J.: LS3MIP (v1.0) contribution to CMIP6: the Land Surface, Snow and Soil moisture Model Intercomparison Project – aims, setup and expected outcome, Geosci. Model Dev., 9, 2809–2832, <a href="https://doi.org/10.5194/gmd-9-2809-2016" target="_blank">https://doi.org/10.5194/gmd-9-2809-2016</a>, 2016.
</mixed-citation></ref-html>
<ref-html id="bib1.bib109"><label>109</label><mixed-citation>
Vander Linden, M.: Code for fig 3 of <a href="https://doi.org/10.5194/gmd-2019-125" target="_blank">https://doi.org/10.5194/gmd-2019-125</a>, available at: <a href="https://github.com/mavdlind/GMD" target="_blank"/>, last access: 14 February 2020.
</mixed-citation></ref-html>
<ref-html id="bib1.bib110"><label>110</label><mixed-citation>
Vavrus, S., Ruddiman, W. F., and Kutzbach, J. E.: Climate model tests of the
anthropogenic influence on greenhouse-induced climate change: the role of
early human agriculture, industrialization, and vegetation feedbacks, Quaternary Sci. Rev., 27, 1410–1425, 2008.
</mixed-citation></ref-html>
<ref-html id="bib1.bib111"><label>111</label><mixed-citation>
Veal, R.: Wood and charcoal for Rome: towards an understanding of ancient regional fuel economics, in: Rural communities in a globalizing economy: new perspectives on the economic integration of Roman Italy, edited by: de Haas, T. and Gijs, T., Brill, New York, Leiden, 388–406, 2017.
</mixed-citation></ref-html>
<ref-html id="bib1.bib112"><label>112</label><mixed-citation>
Viau, A. E. and Gajewski, K.: Reconstructing millennial, regional paleoclimates of boreal Canada during the Holocene, J. Climate, 22, 316–330,
2009.
</mixed-citation></ref-html>
<ref-html id="bib1.bib113"><label>113</label><mixed-citation>
Viau, A. E., Gajewski, K., Sawada, M., and Fines, P.: Mean-continental July temperature variability in North America during the past 14,000 years, J. Geophys. Res.-Atmos., 111, D09102, <a href="https://doi.org/10.1029/2005JD006031" target="_blank">https://doi.org/10.1029/2005JD006031</a>, 2006.
</mixed-citation></ref-html>
<ref-html id="bib1.bib114"><label>114</label><mixed-citation>
Weiberg, E., Hughes, R. E., Finné, M., Bonnier, A., and Kaplan, J. O.:
Mediterranean land use systems from prehistory to antiquity: a case study
from Peloponnese (Greece), J. Land Use Sci., 14, 1–20, <a href="https://doi.org/10.1080/1747423x.2019.1639836" target="_blank">https://doi.org/10.1080/1747423x.2019.1639836</a>, 2019.

</mixed-citation></ref-html>
<ref-html id="bib1.bib115"><label>115</label><mixed-citation>
Whitehouse, N., Schulting, R. J., McClatchie, M., Barratt, P., LcLaughlin, T. R., Bogaard, A., Colledge, S., Marchant, R., Gaffrey, J., and Bunting, M. J.: Neolithic agriculture on the European western frontier: the boom and bust of early farming in Ireland, J. Arch. Sci., 51, 181–205, <a href="https://doi.org/10.1016/j.jas.2013.08.009" target="_blank">https://doi.org/10.1016/j.jas.2013.08.009</a>, 2014.
</mixed-citation></ref-html>
<ref-html id="bib1.bib116"><label>116</label><mixed-citation>
Williams, A.: The use of summed radiocarbon probability distributions in
archaeology: a review of methods, J. Archaeol. Sci., 39, 578–589,
<a href="https://doi.org/10.1016/j.jas.2011.07.014" target="_blank">https://doi.org/10.1016/j.jas.2011.07.014</a>, 2012.
</mixed-citation></ref-html>
<ref-html id="bib1.bib117"><label>117</label><mixed-citation>
Wilmshurst, J. M., McGlone, M. S., Leathwick, J. R., and Newnham, R. M.: A
pre-deforestation pollen-climate calibration model for New Zealand and
quantitative temperature reconstructions for the past 18&thinsp;000 years BP, J.
Quaternay Sci., 22, 535–547, 2007.
</mixed-citation></ref-html>
<ref-html id="bib1.bib118"><label>118</label><mixed-citation>
Wright, P.: Preservation or destruction of plant remains by carbonization, J. Arch. Sci., 30, 577–583, <a href="https://doi.org/10.1016/S0305-4403(02)00203-0" target="_blank">https://doi.org/10.1016/S0305-4403(02)00203-0</a>, 2003.
</mixed-citation></ref-html>
<ref-html id="bib1.bib119"><label>119</label><mixed-citation>
Zahid, H. J., Robinson, E., and Kelly, R. L.: Agriculture, population growth,
and statistical analysis of the radiocarbon record, P. Natl. Acad. Sci. USA,
113, 931–935, <a href="https://doi.org/10.1073/pnas.1517650112" target="_blank">https://doi.org/10.1073/pnas.1517650112</a>, 2016.
</mixed-citation></ref-html>
<ref-html id="bib1.bib120"><label>120</label><mixed-citation>
Zanon, M., Davis, B. A. S., Marquer, L., Brewer, S., and Kaplan, J. O.: European forest cover during the past 12,000 years: a palynological reconstruction based on modern analogs and remote sensing, Front. Plant
Sci., 9, 253, <a href="https://doi.org/10.3389/fpls.2018.00253" target="_blank">https://doi.org/10.3389/fpls.2018.00253</a>, 2018.
</mixed-citation></ref-html>
<ref-html id="bib1.bib121"><label>121</label><mixed-citation>
Zimmermann, A., Wendt, K. P., and Hilpert, J.: Landscape archaeology in
central Europe, Proc. Prehist. Soc., 75, 1–53, <a href="https://doi.org/10.1017/S0079497X00000281" target="_blank">https://doi.org/10.1017/S0079497X00000281</a>, 2009.
</mixed-citation></ref-html>
<ref-html id="bib1.bib122"><label>122</label><mixed-citation>
Zohary, D., Hopf, M., and Weiss, E.: Domestication of Plants in the Old World: The Origin and Spread of Domesticated Plants in South-west Asia,
Europe, and the Mediterranean Basin, 4th Edn., Oxford University Press, Oxford, 2012.
</mixed-citation></ref-html>--></article>
